Plate (animal anatomy) in the context of Pteraspis

In zoology, a scale (Ancient Greek: λεπίς, romanized: lepís; Latin: squāma) is a small rigid plate made out of keratin that grows out of vertebrate animals' skin to provide protection. In lepidopterans (butterflies and moths), scales are plates on the surface of the insect wing, made out of chitin instead of keratin, and provide coloration. Scales are quite common and have evolved multiple times through convergent evolution, with varying structure and function.

Scales are generally classified as part of an organism's integumentary system. There are various types of scales according to the shape and class of an animal.

Placoderms (from Ancient Greek πλάξ [plax, plakos] 'plate' and δέρμα [derma] 'skin') are vertebrate animals of the class Placodermi, an extinct group of prehistoric fish known from Paleozoic fossils during the Silurian and the Devonian periods. While their endoskeletons are mainly cartilaginous, their head and thorax were covered by articulated armoured plates (hence the name), and the rest of the body was scaled or naked depending on the species.

Placoderms were among the first jawed fish (their jaws likely evolved from the first pair of gill arches), as well as the first vertebrates to have true teeth. They were also the first fish clade to develop pelvic fins, the second set of paired fins and the homologous precursor to hindlimbs in tetrapods. 380-million-year-old fossils of three other genera, Incisoscutum, Materpiscis and Austroptyctodus, represent the oldest known examples of live birth.