Phylum in the context of Marine sponge

Biodiversity is the variability of life on Earth. It can be measured on various levels, for example, genetic variability, species diversity, ecosystem diversity and phylogenetic diversity. Diversity is not distributed evenly on Earth—it is greater in the tropics as a result of the warm climate and high primary productivity in the region near the equator. Tropical forest ecosystems cover less than one-fifth of Earth's terrestrial area and contain about 50% of the world's species. There are latitudinal gradients in species diversity for both marine and terrestrial taxa.

Since life began on Earth, six major mass extinctions and several minor events have led to large and sudden drops in biodiversity. The Phanerozoic aeon (the last 540 million years) marked a rapid growth in biodiversity via the Cambrian explosion. In this period, the majority of multicellular phyla first appeared. The next 400 million years included repeated, massive biodiversity losses. Those events have been classified as mass extinction events. In the Carboniferous, rainforest collapse may have led to a great loss of plant and animal life. The Permian–Triassic extinction event, 251 million years ago, was the worst; vertebrate recovery took 30 million years.

Archaea (/ɑːrˈkiːə/ ar-KEE-ə) is a domain of organisms. Traditionally, Archaea included only its prokaryotic members, but has since been found to be paraphyletic, as eukaryotes are known to have evolved from archaea. Even though the domain Archaea cladistically includes eukaryotes, the term archaea (sing. archaeon /ɑːrˈkiːɒn/ ar-KEE-on; from Ancient Greek ἀρχαῖον arkhaîon 'ancient') in English still generally refers specifically to prokaryotic members of Archaea. Archaea were initially classified as bacteria, receiving the name archaebacteria (/ˌɑːrkibækˈtɪəriə/, in the Archaebacteria kingdom), but this term has fallen out of use. Archaeal cells have unique properties separating them from Bacteria and Eukaryota, including: cell membranes made of ether-linked lipids; metabolisms such as methanogenesis; and a unique motility structure known as an archaellum. Archaea are further divided into multiple recognized phyla. Classification is difficult because most have not been isolated in a laboratory and have been detected only by their gene sequences in environmental samples. It is unknown if they can produce endospores.

Archaea are often similar to bacteria in size and shape, although a few have very different shapes, such as the flat, square cells of Haloquadratum walsbyi. Despite this, archaea possess genes and several metabolic pathways that are more closely related to those of eukaryotes, notably for the enzymes involved in transcription and translation. Other aspects of archaeal biochemistry are unique, such as their reliance on ether lipids in their cell membranes, including archaeols. Archaea use more diverse energy sources than eukaryotes, ranging from organic compounds such as sugars, to ammonia, metal ions or even hydrogen gas. The salt-tolerant Halobacteria use sunlight as an energy source, and other species of archaea fix carbon (autotrophy), but unlike cyanobacteria, no known species of archaea does both. Archaea reproduce asexually by binary fission, fragmentation, or budding; unlike bacteria, no known species of Archaea form endospores. The first observed archaea were extremophiles, living in extreme environments such as hot springs and salt lakes with no other organisms. Improved molecular detection tools led to the discovery of archaea in almost every habitat, including soil, oceans, and marshlands. Archaea are particularly numerous in the oceans, and the archaea in plankton may be one of the most abundant groups of organisms on the planet.

The eukaryotes (/juːˈkærioʊts, -əts/) are the domain of Eukaryota or Eukarya, organisms whose cells have a membrane-bound nucleus. All animals, plants, fungi, seaweeds, and many unicellular organisms are eukaryotes. They constitute a major group of life forms alongside the two groups of prokaryotes: the Bacteria and the Archaea. Eukaryotes represent a small minority of the number of organisms, but given their generally much larger size, their collective global biomass is much larger than that of prokaryotes.

The eukaryotes emerged within the archaeal phylum Promethearchaeota. Ignoring mitochondrial DNA (which is bacterial rather than archaeal), this would imply only two domains of life, Bacteria and Archaea, with eukaryotes incorporated among the Archaea. Eukaryotes first emerged during the Paleoproterozoic, likely as flagellated cells. The leading evolutionary theory is they were created by symbiogenesis between an anaerobic Promethearchaeota archaeon and an aerobic proteobacterium, which formed the mitochondria. A second episode of symbiogenesis with a cyanobacterium created the plants, with chloroplasts.

The Cambrian explosion (also known as Cambrian radiation or Cambrian diversification) is an interval of time beginning approximately 538.8 million years ago in the Cambrian period of the early Paleozoic, when a sudden radiation of complex life occurred and practically all major animal phyla started appearing in the fossil record. It lasted for about 13 to 25 million years and resulted in the divergence of most modern metazoan phyla. The event was accompanied by major diversification in other groups of organisms as well.

Before early Cambrian diversification, most organisms were relatively simple, composed of individual cells or small multicellular organisms, occasionally organized into colonies. As the rate of diversification subsequently accelerated, the variety of life became much more complex and began to resemble that of today. Almost all present-day animal phyla appeared during this period, including the earliest chordates.

Animals are multicellular, eukaryotic organisms comprising the biological kingdom Animalia (/ˌænɪˈmeɪliə/). With few exceptions, animals consume organic material, breathe oxygen, have myocytes and are able to move, can reproduce sexually, and grow from a hollow sphere of cells, the blastula, during embryonic development. Animals form a clade, meaning that they arose from a single common ancestor. Over 1.5 million living animal species have been described, of which around 1.05 million are insects, over 85,000 are molluscs, and around 65,000 are vertebrates. It has been estimated there are as many as 7.77 million animal species on Earth. Animal body lengths range from 8.5 μm (0.00033 in) to 33.6 m (110 ft). They have complex ecologies and interactions with each other and their environments, forming intricate food webs. The scientific study of animals is known as zoology, and the study of animal behaviour is known as ethology.

The animal kingdom is divided into five major clades, namely Porifera, Ctenophora, Placozoa, Cnidaria and Bilateria. Most living animal species belong to the clade Bilateria, a highly proliferative clade whose members have a bilaterally symmetric and significantly cephalised body plan, and the vast majority of bilaterians belong to two large clades: the protostomes, which includes organisms such as arthropods, molluscs, flatworms, annelids and nematodes; and the deuterostomes, which include echinoderms, hemichordates and chordates, the latter of which contains the vertebrates. The much smaller basal phylum Xenacoelomorpha have an uncertain position within Bilateria.

Mollusca is a phylum of protostomic invertebrate animals, whose members are known as molluscs or mollusks (/ˈmɒləsks/). Around 76,000 extant species of molluscs are recognized, making it the second-largest animal phylum after Arthropoda. The number of additional fossil species is estimated between 60,000 and 100,000, and the proportion of undescribed species is very high. Many taxa remain poorly studied.

Molluscs are the largest marine phylum, comprising about 23% of all the named marine organisms. They are highly diverse, not just in size and anatomical structure, but also in behaviour and habitat, as numerous groups are freshwater and even terrestrial species. The phylum is typically divided into 7 or 8 taxonomic classes, of which two are entirely extinct. Cephalopod molluscs, such as squid, cuttlefish, and octopuses, are among the most neurologically advanced of all invertebrates—and either the giant squid or the colossal squid is the largest known extant invertebrate species. The gastropods (snails, slugs and abalone) are by far the most diverse class and account for 80% of the total classified molluscan species.

A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of colonies of coral polyps held together by calcium carbonate. Most coral reefs are built from stony corals, whose polyps cluster in groups.

Coral belongs to the class Anthozoa in the animal phylum Cnidaria, which includes sea anemones and jellyfish. Unlike sea anemones, corals secrete hard carbonate exoskeletons that support and protect the coral. Most reefs grow best in warm, shallow, clear, sunny and agitated water. Coral reefs first appeared 485 million years ago, at the dawn of the Early Ordovician, displacing the microbial and sponge reefs of the Cambrian.

In biology, taxonomy (from Ancient Greek τάξις (taxis) 'arrangement' and -νομία (-nomia) 'method') is the scientific study of naming, defining (circumscribing) and classifying groups of biological organisms based on shared characteristics. Organisms are grouped into taxa (singular: taxon), and these groups are given a taxonomic rank; groups of a given rank can be aggregated to form a more inclusive group of higher rank, thus creating a taxonomic hierarchy. The principal ranks in modern use are domain, kingdom, phylum (division is sometimes used in botany in place of phylum), class, order, family, genus, and species. The Swedish botanist Carl Linnaeus is regarded as the founder of the current system of taxonomy, having developed a ranked system known as Linnaean taxonomy for categorizing organisms.

With advances in the theory, data and analytical technology of biological systematics, the Linnaean system has transformed into a system of modern biological classification intended to reflect the evolutionary relationships among organisms, both living and extinct.

Orthogenesis, also known as orthogenetic evolution, progressive evolution, evolutionary progress, or progressionism, is an obsolete biological hypothesis that organisms have an innate tendency to evolve in a definite direction towards some goal (teleology) due to some internal mechanism or "driving force". According to the theory, the largest-scale trends in evolution have an absolute goal such as increasing biological complexity. Prominent historical figures who have championed some form of evolutionary progress include Jean-Baptiste Lamarck, Pierre Teilhard de Chardin, and Henri Bergson.

The term orthogenesis was introduced by Wilhelm Haacke in 1893 and popularized by Theodor Eimer five years later. Proponents of orthogenesis had rejected the theory of natural selection as the organizing mechanism in evolution for a rectilinear (straight-line) model of directed evolution. With the emergence of the modern synthesis, in which genetics was integrated with evolution, orthogenesis and other alternatives to Darwinism were largely abandoned by biologists, but the notion that evolution represents progress is still widely shared; modern supporters include E. O. Wilson and Simon Conway Morris. The evolutionary biologist Ernst Mayr made the term effectively taboo in the journal Nature in 1948, by stating that it implied "some supernatural force". The American paleontologist George Gaylord Simpson (1953) attacked orthogenesis, linking it with vitalism by describing it as "the mysterious inner force". Despite this, many museum displays and textbook illustrations continue to give the impression that evolution is directed.

The nematodes (/ˈnɛmətoʊdz/ NEM-ə-tohdz or NEEM-; Ancient Greek: Νηματώδη; Latin: Nematoda), roundworms or eelworms constitute the phylum Nematoda. Species in the phylum inhabit a broad range of environments. Most species are free-living, feeding on microorganisms, but many are parasitic. Parasitic worms (helminths) are the cause of soil-transmitted helminthiases.

They are classified along with arthropods, tardigrades and other moulting animals in the clade Ecdysozoa. Unlike the flatworms, nematodes have a tubular digestive system, with openings at both ends. Like tardigrades, they have a reduced number of Hox genes, but their sister phylum Nematomorpha has kept the ancestral protostome Hox genotype, which shows that the reduction has occurred within the nematode phylum.

Tardigrades (/ˈtɑːrdɪɡreɪdz/ ), known colloquially as water bears or moss piglets, are a phylum of eight-legged segmented micro-animals. They were first described by the German zoologist Johann August Ephraim Goeze in 1773, who called them Kleiner Wasserbär 'little water bear'. In 1776, the Italian biologist Lazzaro Spallanzani named them Tardigrada, which means 'slow walkers'.

They live in diverse regions of Earth's biosphere – mountaintops, the deep sea, tropical rainforests, and the Antarctic. Tardigrades are among the most resilient animals known, with individual species able to survive extreme conditions – such as exposure to extreme temperatures, extreme pressures (both high and low), air deprivation, radiation, dehydration, and starvation – that would quickly kill most other forms of life. Tardigrades have survived exposure to outer space.

Mosses are small, non-vascular flowerless plants in the taxonomic division Bryophyta (/braɪˈɒfətə/, /ˌbraɪ.əˈfaɪtə/) sensu stricto. Bryophyta (sensu lato, Schimp. 1879) may also refer to the parent group bryophytes, which comprise liverworts, mosses, and hornworts. Mosses typically form dense green clumps or mats, often in damp or shady locations. The individual plants are usually composed of simple leaves that are generally only one cell thick, attached to a stem that may be branched or unbranched and has only a limited role in conducting water and nutrients. Although some species have conducting tissues, these are generally poorly developed and structurally different from similar tissue found in vascular plants. Mosses do not have seeds and after fertilisation develop sporophytes with unbranched stalks topped with single capsules containing spores. They are typically 0.2–10 cm (0.1–3.9 in) tall, though some species are much larger. Dawsonia superba, the tallest moss in the world, can grow to 60 cm (24 in) in height. There are approximately 12,000 species.

Mosses are commonly confused with liverworts, hornworts and lichens. Although often described as non-vascular plants, many mosses have advanced vascular systems. Like liverworts and hornworts, the haploid gametophyte generation of mosses is the dominant phase of the life cycle. This contrasts with the pattern in all vascular plants (seed plants and pteridophytes), where the diploid sporophyte generation is dominant. Lichens may superficially resemble mosses, and sometimes have common names that include the word "moss" (e.g., "reindeer moss" or "Iceland moss"), but they are fungal symbioses and not related to mosses.

Cyanobacteria (/saɪˌænoʊbækˈtɪəriə/ sy-AN-oh-bak-TEER-ee-ə) are a group of autotrophic gram-negative bacteria of the phylum Cyanobacteriota that can obtain biological energy via oxygenic photosynthesis. The name "cyanobacteria" (from Ancient Greek κύανος (kúanos) 'blue') refers to their bluish green (cyan) color, which forms the basis of cyanobacteria's informal common name, blue-green algae.

Cyanobacteria are probably the most numerous taxon to have ever existed on Earth and the first organisms known to have produced oxygen, having appeared in the middle Archean eon and apparently originated in a freshwater or terrestrial environment. Their photopigments can absorb the red- and blue-spectrum frequencies of sunlight (thus reflecting a greenish color) to split water molecules into hydrogen ions and oxygen. The hydrogen ions are used to react with carbon dioxide to produce complex organic compounds such as carbohydrates (a process known as carbon fixation), and the oxygen is released as a byproduct. By continuously producing and releasing oxygen over billions of years, cyanobacteria are thought to have converted the early Earth's anoxic, weakly reducing prebiotic atmosphere, into an oxidizing one with free gaseous oxygen (which previously would have been immediately removed by various surface reductants), resulting in the Great Oxidation Event and the "rusting of the Earth" during the early Proterozoic, dramatically changing the composition of life forms on Earth. The subsequent adaptation of early single-celled organisms to survive in oxygenous environments likely led to endosymbiosis between anaerobes and aerobes, and hence the evolution of eukaryotes during the Paleoproterozoic.

Conifers (/ˈkɒnɪfər/) are a group of seed plants, a subset of gymnosperms. They are mainly evergreen trees with a regular branching pattern, reproducing with male and female cones, usually on the same tree. They are wind-pollinated and the seeds are usually dispersed by the wind. Scientifically, they make up the division Pinophyta, also known as Coniferae. All extant conifers except for the Gnetophytes are perennial woody plants with secondary growth. There are over 600 living species.

Conifers first appear in the fossil record over 300 million years ago in the Carboniferous. They became dominant land plants in the Mesozoic, until flowering plants took over many ecosystems in the Cretaceous. Many conifers today are relict species, surviving in a small part of their former ranges. Such relicts include Wollemia, known only from a small area of Australia, and Metasequoia glyptostroboides, known from Cretaceous fossils and surviving in a small area of China.

The euphyllophytes are a clade of plants within the tracheophytes (the vascular plants). The group may be treated as an unranked clade, a division under the name Euphyllophyta or a subdivision under the name Euphyllophytina. The euphyllophytes are characterized by the possession of true leaves ("megaphylls"), and comprise one of two major lineages of extant vascular plants. As shown in the cladogram below, the euphyllophytes have a sister relationship to the lycopodiophytes or lycopsids. Unlike the lycopodiophytes, which consist of relatively few presently living or extant taxa, the euphyllophytes comprise the vast majority of vascular plant lineages that have evolved since both groups shared a common ancestor more than 400 million years ago. The euphyllophytes consist of two lineages, the spermatophytes or seed plants such as flowering plants (angiosperms) and gymnosperms (conifers and related groups), and the Polypodiophytes or ferns, as well as a number of extinct fossil groups.

The division of the extant tracheophytes into three monophyletic lineages is supported in multiple molecular studies. Other researchers argue that phylogenies based solely on molecular data without the inclusion of carefully evaluated fossil data based on whole plant reconstructions, do not necessarily completely and accurately resolve the evolutionary history of groups like the euphyllophytes.