Phylogenetic nomenclature in the context of "Heteroptera"

In biological taxonomy, taxonomic rank (which some authors prefer to call nomenclatural rank because ranking is part of nomenclature rather than taxonomy proper, according to some definitions of these terms) is the relative or absolute level of a group of organisms (a taxon) in a hierarchy that reflects evolutionary relationships. Thus, the most inclusive clades (such as Eukarya and Animalia) have the highest ranks, whereas the least inclusive ones (such as Homo sapiens or Bufo bufo) have the lowest ranks. Ranks can be either relative and be denoted by an indented taxonomy in which the level of indentation reflects the rank, or absolute, in which various terms, such as species, genus, family, order, class, phylum, kingdom, and domain designate rank. This page emphasizes absolute ranks and the rank-based codes (the Zoological Code, the Botanical Code, the Code for Cultivated Plants, the Prokaryotic Code, and the Code for Viruses) require them. However, absolute ranks are not required in all nomenclatural systems for taxonomists; for instance, the PhyloCode, the code of phylogenetic nomenclature, does not require absolute ranks.

Taxa are hierarchical groups of organisms, and their ranks describes their position in this hierarchy. High-ranking taxa (e.g. those considered to be domains or kingdoms, for instance) include more sub-taxa than low-ranking taxa (e.g. those considered genera, species or subspecies). The rank of these taxa reflects inheritance of traits or molecular features from common ancestors. The name of any species and genus are basic; which means that to identify a particular organism, it is usually not necessary to specify names at ranks other than these first two, within a set of taxa covered by a given rank-based code. However, this is not true globally because most rank-based codes are independent from each other, so there are many inter-code homonyms (the same name used for different organisms, often for an animal and for a taxon covered by the botanical code). For this reason, attempts were made at creating a BioCode that would regulate all taxon names, but this attempt has so far failed because of firmly entrenched traditions in each community.

In biology, the taxon (back-formation from taxonomy; pl.: taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and given a particular ranking, especially if and when it is accepted or becomes established. It is very common, however, for taxonomists to remain at odds over what belongs to a taxon and the criteria used for inclusion, especially in the context of rank-based ("Linnaean") nomenclature (much less so under phylogenetic nomenclature). If a taxon is given a formal scientific name, its use is then governed by one of the nomenclature codes specifying which scientific name is correct for a particular grouping.

Initial attempts at classifying and ordering organisms (plants and animals) were presumably set forth in prehistoric times by hunter-gatherers, as suggested by the fairly sophisticated folk taxonomies. Much later, Aristotle and, later still, European scientists, like Magnol, Tournefort and Carl Linnaeus's system in Systema Naturae, 10th edition (1758),, as well as an unpublished work by Bernard and Antoine Laurent de Jussieu, contributed to this field. The idea of a unit-based system of biological classification was first made widely available in 1805 in the introduction to Jean-Baptiste Lamarck's Flore françoise, and Augustin Pyramus de Candolle's Principes élémentaires de botanique. Lamarck set out a system for the "natural classification" of plants. Since then, systematists continue to construct accurate classifications encompassing the diversity of life; today, a "good" or "useful" taxon is commonly taken to be one that reflects evolutionary relationships.

SAR is a highly diverse clade of eukaryotes, often considered a supergroup, that includes stramenopiles (heterokonts), alveolates, and rhizarians. It is a node-based taxon (under the Sar name), including all descendants of the three groups' last common ancestor, and comprises most of the now-rejected Chromalveolata. Their sister group might be telonemids, with which they would make up the TSAR (sometimes called T-SAR) clade, but this grouping has been challenged. Harosa is used synonymously with TSAR by Cavalier-Smith in 2022.

The International Code of Phylogenetic Nomenclature, known as the PhyloCode for short, is a formal set of rules governing phylogenetic nomenclature. Its current version is specifically designed to regulate the naming of clades, leaving the governance of species names up to the rank-based nomenclature codes (ICN, ICNCP, ICNP, ICZN, ICVCN).

The PhyloCode is associated with the International Society for Phylogenetic Nomenclature (ISPN). The companion volume, Phylonyms, establishes 300 taxon names under PhyloCode, serving as examples for those unfamiliar with the code. RegNum is an associated online database for registered clade names.