Mesozoic in the context of Cambrian explosion

Hvar (pronounced [xvâːr]; Chakavian: Hvor or For; Greek: Φάρος, romanized: Pharos; Latin: Pharia; Italian: Lesina) is a Croatian island in the Adriatic Sea, located off the Dalmatian coast, lying between the islands of Brač, Vis and Korčula. Approximately 68 kilometres (42.25 mi) long, with a high east–west ridge of Mesozoic limestone and dolomite, the island of Hvar is unusual in the area for having a large fertile coastal plain, and fresh water springs. Its hillsides are covered in pine forests, with vineyards, olive groves, fruit orchards and lavender fields in the agricultural areas. The climate is characterized by mild winters, and warm summers with many hours of sunshine. The island has 10,678 residents according to the 2021 census, making it the fourth most populated of the Croatian islands.

The Jurassic (/dʒʊˈræsɪk/ juurr-ASS-ik) is a geologic period and stratigraphic system that spanned from the end of the Triassic Period 201.4 Ma (million years ago) to the beginning of the Cretaceous Period, 143.1 Ma. The Jurassic constitutes the second and middle period of the Mesozoic Era as well as the eighth period of the Phanerozoic Eon and is named after the Jura Mountains, where limestone strata from the period were first identified.

The start of the Jurassic was marked by the major Triassic–Jurassic extinction event, associated with the eruption of the Central Atlantic Magmatic Province (CAMP). The beginning of the Toarcian Age started around 183 Ma and is marked by the Toarcian Oceanic Anoxic Event, a global episode of oceanic anoxia, ocean acidification, and elevated global temperatures associated with extinctions, likely caused by the eruption of the Karoo-Ferrar large igneous provinces. The end of the Jurassic, however, has no clear, definitive boundary with the Cretaceous and is the only boundary between geological periods to remain formally undefined.

The Moenave Formation is a Mesozoic geologic formation, in the Glen Canyon Group. It is found in Utah and Arizona.

The Moenave was deposited on an erosion surface on the Chinle Formation following an early Jurassic uplift and unconformity that represents about ten million years of missing sedimentation. Periodic incursions of shallow seas from the north during the Jurassic flooded parts of Wyoming, Montana, and a northeast–southwest trending trough on the Utah/Idaho border. The Moenave was deposited in a variety of river, lake, and flood-plain environments, near the ancient Lake Dixie.

Petroleum, also known as crude oil or simply oil, is a naturally occurring, yellowish-black liquid chemical mixture found in geological formations, consisting mainly of hydrocarbons. The term petroleum refers both to naturally occurring unprocessed crude oil, as well as to petroleum products that consist of refined crude oil.

Petroleum is a fossil fuel formed over millions of years from anaerobic decay of organic materials from buried prehistoric organisms, particularly planktons and algae. It is estimated that 70% of the world's oil deposits were formed during the Mesozoic, 20% were formed in the Cenozoic, and only 10% were formed in the Paleozoic. Conventional reserves of petroleum are primarily recovered by drilling, which is done after a study of the relevant structural geology, analysis of the sedimentary basin, and characterization of the petroleum reservoir. There are also unconventional reserves such as oil sands and oil shale which are recovered by other means such as fracking.

Maastricht (/ˈmɑːstrɪxt/ MAH-strikht, US also /mɑːˈstrɪxt/ mah-STRIKHT, Dutch: [maːˈstrɪxt] ; Limburgish: Mestreech [məˈstʀeːx]) is a city and a municipality in the southeastern Netherlands. It is the capital and largest city of the province of Limburg. Maastricht is located on both sides of the Meuse (Dutch: Maas), at the point where the river is joined by the Jeker. Mount Saint Peter (Sint-Pietersberg) is largely situated within the city's municipal borders. Maastricht is adjacent to the border with Belgium and is part of the Meuse-Rhine Euroregion, an international metropolis with a population of about 3.9 million, which includes the nearby German and Belgian cities of Aachen, Liège, and Hasselt.

Maastricht developed from a Roman settlement (Trajectum ad Mosam) to a medieval river trade and religious centre. In the 16th century it became a garrison town and in the 19th century an early industrial centre. Today, the city is a thriving cultural and regional hub. It became well known through the Maastricht Treaty and as the birthplace of the euro. Maastricht has 1,677 national heritage buildings (rijksmonumenten), the second highest number in the Netherlands, after Amsterdam. The city is visited by tourists for shopping and recreation, and has a large international student population. The last stage of the Cretaceous period and the Mesozoic era, the Maastrichtian, is named after this city, at the end of which was the Cretaceous–Paleogene extinction event, which resulted in the extinction of the non-avian dinosaurs.

The Cenozoic Era (also known as Caenozoic, Kainozoic, or Neozoic Era; /ˌsiːnəˈzoʊ.ɪk, ˌsɛn-/; SEE-nə-ZOH-ik, SEN-ə-; lit. 'new life') is Earth's current geological era, representing the last 66 million years of Earth's history. It is characterized by the dominance of mammals, insects, birds and angiosperms (flowering plants). It is the latest of three geological eras of the Phanerozoic Eon, preceded by the Mesozoic and Paleozoic. The Cenozoic started with the Cretaceous–Paleogene extinction event, when many species, including the non-avian dinosaurs, became extinct in an event attributed by most experts to the impact of a large asteroid or other celestial body, the Chicxulub impactor.

The Cenozoic is also known as the Age of Mammals because the terrestrial animals that dominated both hemispheres were mammals – the eutherians (placentals) in the Northern Hemisphere and the metatherians (marsupials, now mainly restricted to Australia and to some extent South America) in the Southern Hemisphere. The extinction of many groups allowed mammals and birds to greatly diversify so that large mammals and birds dominated life on Earth. The continents also moved into their current positions during this era.

Conifers (/ˈkɒnɪfər/) are a group of seed plants, a subset of gymnosperms. They are mainly evergreen trees with a regular branching pattern, reproducing with male and female cones, usually on the same tree. They are wind-pollinated and the seeds are usually dispersed by the wind. Scientifically, they make up the division Pinophyta, also known as Coniferae. All extant conifers except for the Gnetophytes are perennial woody plants with secondary growth. There are over 600 living species.

Conifers first appear in the fossil record over 300 million years ago in the Carboniferous. They became dominant land plants in the Mesozoic, until flowering plants took over many ecosystems in the Cretaceous. Many conifers today are relict species, surviving in a small part of their former ranges. Such relicts include Wollemia, known only from a small area of Australia, and Metasequoia glyptostroboides, known from Cretaceous fossils and surviving in a small area of China.

The Triassic (/traɪˈæsɪk/; sometimes symbolized as 🝈) is a geologic period and a stratigraphic system that spans 50.5 million years from the end of the Permian Period 251.902 Ma (million years ago) to the beginning of the Jurassic Period 201.4 Ma. The Triassic Period is the first and shortest geologic period of the Mesozoic Era, and the seventh period of the Phanerozoic Eon. The start and the end of the Triassic Period featured major extinction events.

Chronologically, the Triassic Period is divided into three epochs: (i) the Early Triassic, (ii) the Middle Triassic, and (iii) the Late Triassic. The Triassic Period began after the Permian–Triassic extinction event that much reduced the biosphere of planet Earth. The fossil record of the Triassic Period presents three categories of organisms: (i) animals that survived the Permian–Triassic extinction event, (ii) new animals that briefly flourished in the Triassic biosphere, and (iii) new animals that evolved and dominated the Mesozoic Era. Reptiles, especially archosaurs, were the chief terrestrial vertebrates during this time. A specialized group of archosaurs, called dinosaurs, first appeared in the Late Triassic but did not become dominant until the succeeding Jurassic Period. Archosaurs that became dominant in this period were primarily pseudosuchians, relatives and ancestors of modern crocodilians, while some archosaurs specialized in flight, the first time among vertebrates, becoming the pterosaurs. Therapsids, the dominant vertebrates of the preceding Permian period, saw a brief surge in diversification in the Triassic, with dicynodonts and cynodonts quickly becoming dominant, but they declined throughout the period with the majority becoming extinct by the end. However, the first stem-group mammals (mammaliamorphs), themselves a specialized subgroup of cynodonts, appeared during the Triassic and would survive the extinction event, allowing them to radiate during the Jurassic. Amphibians were primarily represented by the temnospondyls, giant aquatic predators that had survived the end-Permian extinction and saw a new burst of diversification in the Triassic, before going extinct by the end; however, early crown-group lissamphibians (including stem-group frogs, salamanders and caecilians) also became more common during the Triassic and survived the extinction event. The earliest known neopterygian fish, including early holosteans and teleosts, appeared near the beginning of the Triassic, and quickly diversified to become among the dominant groups of fish in both freshwater and marine habitats.

Crocodilia (/krɒkəˈdɪliə/) is an order of semiaquatic, predatory reptiles that are known as crocodilians. They appeared 83.5 million years ago in the Late Cretaceous period (Campanian stage) and are the closest living relatives of birds, as the two groups are the only known survivors of the Archosauria. Members of the crocodilian total group, the clade Pseudosuchia, appeared about 250 million years ago in the Early Triassic period, and diversified during the Mesozoic era. The order includes the true crocodiles (family Crocodylidae), the alligators and caimans (family Alligatoridae), and the gharial and false gharial (family Gavialidae). Although the term "crocodiles" is sometimes used to refer to all of these families, the term "crocodilians" is less ambiguous.

Extant crocodilians have flat heads with long snouts and tails that are compressed on the sides, with their eyes, ears, and nostrils at the top of the head. Alligators and caimans tend to have broader U-shaped jaws that, when closed, show only the upper teeth, whereas crocodiles usually have narrower V-shaped jaws with both rows of teeth visible when closed. Gharials have extremely slender, elongated jaws. The teeth are conical and peg-like, and the bite is powerful. All crocodilians are good swimmers and can move on land in a "high walk" position, traveling with their legs erect rather than sprawling. Crocodilians have thick skin covered in non-overlapping scales and, like birds, have a four-chambered heart and lungs with unidirectional airflow.

Archaeopteryx (/ˌɑːrkiːˈɒptərɪks/ ; lit. 'ancient wing'), sometimes referred to by its German name, "Urvogel " (lit. 'Primeval Bird') is a genus of bird-like dinosaurs. The genus name derives from the Ancient Greek ἀρχαῖος (archaîos), meaning 'ancient', and πτέρυξ (ptérux), meaning 'feather, wing'. Between the late 19th century and the early 21st century, Archaeopteryx was generally accepted by palaeontologists and popular reference books as the oldest known bird (member of the group Avialae). Older potential avialans have since been identified, including Anchiornis, Xiaotingia, Aurornis, and Baminornis.

Archaeopteryx lived in the Late Jurassic around 150 million years ago, in what is now southern Germany, during a time when Europe was an archipelago of islands in a shallow warm tropical sea, much closer to the equator than it is now. Similar in size to a Eurasian magpie, with the largest individuals possibly attaining the size of a raven, the largest species of Archaeopteryx could grow to about 50 cm (20 in) in length. Despite their small size, broad wings, and inferred ability to fly or glide, Archaeopteryx had more in common with other small Mesozoic dinosaurs than with modern birds. In particular, they shared the following features with the dromaeosaurids and troodontids: jaws with sharp teeth, three fingers with claws, a long bony tail, hyperextensible second toes ("killing claw"), feathers (which also suggest warm-bloodedness), and various features of the skeleton.

The Cretaceous–Paleogene (K–Pg) extinction event, formerly known as the Cretaceous-Tertiary (K–T) extinction event, was a major mass extinction of three-quarters of the plant and animal species on Earth approximately 66 million years ago. The event caused the extinction of all non-avian dinosaurs. Most other tetrapods weighing more than 25 kg (55 lb) also became extinct, with the exception of some ectothermic species such as sea turtles and crocodilians. It marked the end of the Cretaceous period, and with it the Mesozoic era, while heralding the beginning of the current geological era, the Cenozoic Era. In the geologic record, the K–Pg event is marked by a thin layer of sediment called the K–Pg boundary or K–T boundary, which can be found throughout the world in marine and terrestrial rocks. The boundary clay shows unusually high levels of the metal iridium, which is more common in asteroids than in the Earth's crust.

As originally proposed in 1980 by a team of scientists led by Luis Alvarez and his son Walter, it is now generally thought that the K–Pg extinction resulted from the impact of a massive asteroid 10 to 15 km (6 to 9 mi) wide, 66 million years ago, causing the Chicxulub impact crater and devastating the global environment, mainly through a lingering impact winter which halted photosynthesis in plants and plankton. The impact hypothesis, also known as the Alvarez hypothesis, was bolstered by the discovery of the 180 km (112 mi) Chicxulub crater in the Gulf of Mexico's Yucatán Peninsula in the early 1990s. The temporal match between the ejecta layer, and the onset of the extinctions and the agreement of ecological patterns in the fossil record with modeled environmental perturbations (for example, darkness and cooling), lead to the conclusion that the Chicxulub impact triggered the mass extinction. A 2016 drilling project into the Chicxulub peak ring confirmed that the peak ring comprised granite ejected within minutes from deep in the Earth, but contained hardly any gypsum, the usual sulfate-containing sea floor rock in the region: the gypsum would have vaporized and dispersed as an aerosol into the atmosphere, causing longer-term effects on the climate and food chain. In October 2019, researchers proposed the mechanisms of the mass extinction, arguing that the Chicxulub asteroid impact event rapidly acidified the oceans and produced long-lasting effects on the climate.

Pterosaurs are an extinct clade of flying reptiles in the order Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 million to 66 million years ago). Pterosaurs are the earliest vertebrates known to have evolved powered flight. Their wings were formed by a membrane of skin, muscle, and other tissues stretching from the ankles to a dramatically lengthened fourth finger.

Traditionally, pterosaurs were divided into two major types. Basal pterosaurs (also called non-pterodactyloid pterosaurs or 'rhamphorhynchoids') were smaller animals, up to two meter wingspan, with fully toothed jaws and, typically, long tails. Their wide wing membranes probably included and connected the hindlimbs. On the ground, they would have had an awkward sprawling posture due to short metacarpals, but the anatomy of their joints and strong claws would have made them effective climbers, and some may have lived in trees. Basal pterosaurs were insectivores, piscivores or predators of small land vertebrates. Later pterosaurs (pterodactyloids) evolved many sizes, shapes, and lifestyles. Pterodactyloids had narrower wings with free hindlimbs, highly reduced tails, and long necks with large heads. On the ground, they walked well on all four limbs due to long metacarpals with an upright posture, standing plantigrade on the hind feet and folding the wing finger upward to walk on the metacarpals with the three smaller fingers of the hand pointing to the rear. They could take off from the ground, and fossil trackways show that at least some species were able to run, wade, and/or swim. Their jaws had horny beaks, and some groups lacked teeth. Some groups developed elaborate head crests with sexual dimorphism. Since 2010 it is understood that many species, the basal Monofenestrata, were intermediate in build, combining an advanced long skull with long tails.

The Permian–Triassic extinction event, colloquially known as the Great Dying, was an extinction event that occurred approximately 251.9 million years ago (mya), at the boundary between the Permian and Triassic geologic periods, and with them the Paleozoic and Mesozoic eras. It is Earth's most severe known extinction event, with the extinction of 57% of biological families, 62% of genera, 81% of marine species, and 70% of terrestrial vertebrate species. It is also the greatest known mass extinction of insects. It is the greatest of the "Big Five" mass extinctions of the Phanerozoic. There is evidence for one to three distinct pulses, or phases, of extinction.

The scientific consensus is that the main cause of the extinction was the flood basalt volcanic eruptions that created the Siberian Traps, which released sulfur dioxide and carbon dioxide, resulting in euxinia (oxygen-starved, sulfurous oceans), elevated global temperatures,and acidified oceans.The level of atmospheric carbon dioxide rose from around 400 ppm to 2,500 ppm with approximately 3,900 to 12,000 gigatonnes of carbon being added to the ocean-atmosphere system during this period.

Marine reptiles are reptiles which have become secondarily adapted for an aquatic or semiaquatic life in a marine environment. Only about 100 of the 12,000 extant reptile species and subspecies are classed as marine reptiles, including marine iguanas, sea snakes, sea turtles and saltwater crocodiles.

The earliest marine reptile was Mesosaurus (not to be confused with Mosasaurus), which arose in the Permian period of the Paleozoic era. During the Mesozoic era, many groups of reptiles became adapted to life in the seas, including such familiar clades as the ichthyosaurs, plesiosaurs (these two orders were once thought united in the group "Enaliosauria", a classification now cladistically obsolete), mosasaurs, nothosaurs, placodonts, sea turtles, thalattosaurs and thalattosuchians. Most marine reptile groups became extinct at the end of the Cretaceous period, but some still existed during the Cenozoic, most importantly the sea turtles. Other Cenozoic marine reptiles included the bothremydids, palaeophiid snakes, a few choristoderes such as Simoedosaurus and dyrosaurid crocodylomorphs. Various types of marine gavialid crocodilians remained widespread as recently as the Late Miocene.

The Paleozoic (/ˌpæli.əˈzoʊ.ɪk, -i.oʊ-, ˌpeɪ-/ PAL-ee-ə-ZOH-ik, -⁠ee-oh-, PAY-; or Palaeozoic) Era is the first of three geological eras of the Phanerozoic Eon. Beginning 538.8 million years ago (Ma), it succeeds the Neoproterozoic (the last era of the Proterozoic Eon) and ends 251.9 Ma at the start of the Mesozoic Era. The Paleozoic is subdivided into six geologic periods, (from oldest to youngest) Cambrian, Ordovician, Silurian, Devonian, Carboniferous and Permian. Some geological timescales divide the Paleozoic informally into early and late sub-eras: the Early Paleozoic consisting of the Cambrian, Ordovician and Silurian; the Late Paleozoic consisting of the Devonian, Carboniferous and Permian.

The name Paleozoic was first used by Adam Sedgwick (1785–1873) in 1838 to describe the Cambrian and Ordovician periods. It was redefined by John Phillips (1800–1874) in 1840 to cover the Cambrian to Permian periods. It is derived from the Greek palaiós (παλαιός, "old") and zōḗ (ζωή, "life") meaning "ancient life".

Pangaea or Pangea (/pænˈdʒiːə/ pan-JEE-ə) was a supercontinent that existed during the late Paleozoic and early Mesozoic eras. It assembled from the earlier continental units of Gondwana, Euramerica and Siberia during the Carboniferous period approximately 335 million years ago, and began to break apart about 200 million years ago, at the end of the Triassic and beginning of the Jurassic. Pangaea was C-shaped, with the bulk of its mass stretching between Earth's northern and southern polar regions and surrounded by the superocean Panthalassa and the Paleo-Tethys and subsequent Tethys Oceans. Pangaea is the most recent supercontinent to have existed and was the first to be reconstructed by geologists.

Pteridospermatophyta, also called pteridosperms or seed ferns, are a polyphyletic grouping of extinct seed-producing plants. The earliest fossil evidence for plants of this type are the lyginopterids of late Devonian age. They flourished particularly during the Carboniferous and Permian periods. Pteridosperms declined during the Mesozoic Era and had mostly disappeared by the end of the Cretaceous Period, though Komlopteris seem to have survived into Eocene times, based on fossil finds in Tasmania.

With regard to the enduring utility of this division, many palaeobotanists still use the pteridosperm grouping in an informal sense to refer to the seed plants that are not angiosperms, coniferoids (conifers or cordaites), ginkgophytes (ginkgos or czekanowskiales), cycadophytes (cycads or bennettites), or gnetophytes. This is particularly useful for extinct seed plant groups whose systematic relationships remain speculative, as they can be classified as pteridosperms with no invalid implications being made as to their systematic affinities. Also, from a purely curatorial perspective the term pteridosperms is a useful shorthand for describing the fern-like fronds that were probably produced by seed plants, which are commonly found in many Palaeozoic and Mesozoic fossil floras.

Biostratigraphy is the branch of stratigraphy which focuses on correlating and assigning relative ages of rock strata by using the fossil assemblages contained within them. The primary objective of biostratigraphy is correlation, demonstrating that a particular horizon in one geological section represents the same period of time as another horizon at a different section. Fossils within these strata are useful because sediments of the same age can look completely different, due to local variations in the sedimentary environment. For example, one section might have been made up of clays and marls, while another has more chalky limestones. However, if the fossil species recorded are similar, the two sediments are likely to have been laid down around the same time. Ideally these fossils are used to help identify biozones, as they make up the basic biostratigraphy units, and define geological time periods based upon the fossil species found within each section.

Basic concepts of biostratigraphic principles were introduced in the early 1800s. A Danish scientist and bishop by the name of Nicolas Steno was one of the first geologists to recognize that rock layers correlate to the Law of Superposition. With advancements in science and technology, by the 18th century it began to be accepted that fossils were remains left by species that had become extinct, but were then preserved within the rock record. The method was well-established before Charles Darwin explained the mechanism behind it—evolution. Scientists William Smith, George Cuvier, and Alexandre Brongniart came to the conclusion that fossils then indicated a series of chronological events, establishing layers of rock strata as some type of unit, later termed biozone. From here on, scientists began relating the changes in strata and biozones to different geological eras, establishing boundaries and time periods within major faunal changes. By the late 18th century the Cambrian and Carboniferous periods were internationally recognized due to these findings. During the early 20th century, advancements in technology gave scientists the ability to study radioactive decay. Using this methodology, scientists were able to establish geological time, the boundaries of the different eras (Paleozoic, Mesozoic, Cenozoic), as well as Periods (Cambrian, Ordovician, Silurian) through the isotopes found within fossils via radioactive decay. Current 21st century uses of biostratigraphy involve interpretations of age for rock layers, which are primarily used by oil and gas industries for drilling workflows and resource allocations.

Borneo (/ˈbɔːrnioʊ/) is the third-largest island in the world, with an area of 748,168 km (288,869 sq mi), and population of 23,053,723 (2020 national censuses). Situated at the geographic centre of Maritime Southeast Asia, it is one of the Greater Sunda Islands, located north of Java, west of Sulawesi, and east of Sumatra. The island is crossed by the equator, which divides it roughly in half. In Indonesia, the island is also known as Kalimantan, which is also the name of the Indonesian region located on the island.

The geology of Borneo was formed beginning in the Mesozoic. It formed part of Sundaland, a region connected to mainland Asia, until it became isolated by sea level rise at the end of the last ice age. With seven unique ecoregions, including large expanses of some of the oldest tropical rainforests in the world, Borneo is rich in biodiversity and endemic species.

The Tethys Ocean (/ˈtiːθɪs, ˈtɛ-/ TEETH-iss, TETH-; Greek: Τηθύς Tēthús), also called the Tethys Sea or the Neo-Tethys, was a prehistoric ocean during much of the Mesozoic Era and early-mid Cenozoic Era. It was the predecessor to the modern Indian Ocean, the Mediterranean Sea, and the Eurasian inland marine basins (primarily represented today by the Black Sea and Caspian Sea).

During the early Mesozoic, as Pangaea broke up, the designation “Tethys Ocean” refers to the ocean located between the ancient continents of Gondwana and Laurasia. After the opening of the Indian and Atlantic oceans during the Cretaceous Period and the breakup of these continents over the same period, it refers to the ocean bordered by the continents of Africa, Eurasia, India, and Australasia. During the early-mid Cenozoic, the Indian, African, Australian and Arabian plates moved north and collided with the Eurasian plate, which created new borders to the ocean, a land barrier to the flow of currents between the Indian and Mediterranean basins, and the orogenies of the Alpide belt (including the Alps, Himalayas, Zagros, and Caucasus Mountains). All of these geological events, in addition to a drop in sea level from Antarctic glaciation, brought an end to the Tethys as it previously existed, fragmenting it into the Indian Ocean, the Mediterranean Sea, and the Paratethys.