Lake Zumpango in the context of "Toltec"

The Valley of Mexico (Spanish: Valle de México; Nahuatl languages: Anahuac, lit. 'Land Between the Waters'), sometimes also called Basin of Mexico, is a highlands plateau in central Mexico. Surrounded by mountains and volcanoes, the Valley of Mexico was a centre for several pre-Columbian civilizations including Teotihuacan, the Toltec, and the Aztec Empire. The valley used to contain five interconnected lakes called Lake Zumpango, Lake Xaltocan (Nahuatl languages: Xāltocān), Lake Xochimilco, Lake Chalco and the largest, Lake Texcoco, covering about 1,500 square kilometers (580 sq mi) of the valley floor. When the Spaniards arrived in the Valley of Mexico, it had one of the highest population concentrations in the world with about one million people. After the conquest of the Aztec Empire, the Spaniards rebuilt the largest and most dominant city, Mēxihco Tenōchtitlan, renaming it Ciudad de México (Mexico City) and over time began to drain the lakes' waters to control flooding.

The Valley of Mexico is located in the Trans-Mexican Volcanic Belt. The valley contains most of the Mexico City metropolitan area, as well as parts of the State of Mexico, Hidalgo, Tlaxcala, and Puebla. The Basin of Mexico covers approximately 9,600 km (3,700 sq mi) in the NNE-SSW direction with length to width dimensions of approximately 125 km (78 miles) to 75 km (47 miles) The Valley of Mexico can be subdivided into four basins, but the largest and most-studied is the area that contains Mexico City. This section of the valley in particular is colloquially referred to as the "Valley of Mexico".The valley has a minimum elevation of 2,200 meters (7,200 ft) above sea level and is surrounded by mountains and volcanoes that reach elevations of over 5,000 meters (16,000 ft). It is an enclosed valley with no natural outlet for water to flow to the sea although there is a gap to the north where there is a high mesa but no high mountain peaks. Within this vulnerable watershed all the native fishes were extinct by the end of the 20th century. Hydrologically, the valley has three features. The first feature is the lakebeds of five now-extinct lakes, which are located in the southernmost and largest of the four sub-basins. The other two features are piedmont, and the mountainsides that collect the precipitation that eventually flows to the lake area. These last two are found in all four of the sub-basins of the valley. Today, the Valley drains through a series of artificial canals to the Tula River, and eventually the Páruco River and the Gulf of Mexico. Seismic activity is frequent here, and the valley is an earthquake-prone zone.

The axolotl (/ˈæksəlɒtəl/ ; from Classical Nahuatl: āxōlōtl [aːˈʃoːloːtɬ] ) (Ambystoma mexicanum) is a species of paedomorphic mole salamander, meaning they mature without undergoing metamorphosis into the terrestrial adult form; the adults remain fully aquatic with obvious external gills. This trait, although somewhat unusual among the majority of amphibians, is not unique to axolotls. Furthermore, this is apparent as axolotls may be difficult to distinguish from the larval stage of other neotenic adult mole salamanders (Ambystoma spp.), such as the occasionally paedomorphic tiger salamander (A. tigrinum) which are widespread in North America; or with mudpuppies (Necturus spp.), which bear a superficial resemblance to axolotls, but are from a different family of salamanders.

Axolotls originally inhabited a system of interconnected wetlands and lakes in the Mexican highlands; they were known to inhabit the smaller lakes of Xochimilco and Chalco, and are also presumed to have inhabited the larger lakes of Texcoco and Zumpango. These waterways were mostly drained by Spanish settlers after the conquest of the Aztec Empire, leading to the destruction of much of the axolotl's natural habitat, which is now largely occupied by Mexico City. Despite this, they remained abundant enough to form part of the staple in the diet of native Mexica during the colonial era. Today, due to continued urbanization in Mexico City, which causes water pollution in the remaining waterways, as well as the introduction of invasive species such as tilapia and carp, the axolotls are now near extinction. This can be seen as the species has been listed as critically endangered in the wild, with a decreasing population of around 50 to 1,000 adult individuals, by the International Union for Conservation of Nature (IUCN), and is listed under Appendix II of the Convention on International Trade in Endangered Species (CITES).