Hominin in the context of Australopithecus anamensis

Prehistory, also called pre-literary history, is the period of human history between the first known use of stone tools by hominins c. 3.3 million years ago and the beginning of recorded history with the invention of writing systems. The use of symbols, marks, and images appears very early among humans, but the earliest known writing systems appeared c. 5,200 years ago. It took thousands of years for writing systems to be widely adopted, with writing having spread to almost all cultures by the 19th century. The end of prehistory therefore came at different times in different places, and the term is less often used in discussing societies where prehistory ended relatively recently. It is based on an old conception of history that without written records there could be no history. The most common conception today is that history is based on evidence, however the concept of prehistory has not been completely discarded.

In the early Bronze Age, Sumer in Mesopotamia, the Indus Valley Civilisation, and ancient Egypt were the first civilizations to develop their own scripts and keep historical records, with their neighbours following. Most other civilizations reached their end of prehistory during the following Iron Age. The three-age division of prehistory into Stone Age, Bronze Age, and Iron Age remains in use for much of Eurasia and North Africa, but is not generally used in those parts of the world where the working of hard metals arrived abruptly from contact with Eurasian cultures, such as Oceania, Australasia, much of Sub-Saharan Africa, and parts of the Americas. With some exceptions in pre-Columbian civilizations in the Americas, these areas did not develop writing systems before the arrival of Eurasians, so their prehistory reaches into relatively recent periods; for example, 1788 is usually taken as the end of the prehistory of Australia.

Homo (from Latin homō 'human') is a genus of great ape (family Hominidae) that emerged from the early homininian genus Australopithecus, encompassing a single extant species, Homo sapiens (modern humans), along with a number of extinct species (e.g. Homo erectus and Homo neanderthalensis) classified as either ancestral or closely related to modern humans, collectively called archaic humans. Homo, together with the genus Paranthropus, is probably most closely related to the species Australopithecus africanus within Australopithecus. The closest living relatives of Homo are of the hominin genus Pan (chimpanzees and bonobos), with the ancestors of Pan and Homo estimated to have diverged around 5.7–11 million years ago during the Late Miocene.

The oldest member of the genus is Homo habilis, with fossil records of just over 2 million years ago. H. erectus appeared about 2 million years ago and spread throughout Africa (debatably as another species called Homo ergaster) and Eurasia in several migrations. The species was adaptive and successful, and persisted for more than a million years before gradually diverging into new species around 500,000 years ago.

The Levantine corridor is the relatively narrow strip in Western Asia, between the Mediterranean Sea to the northwest and deserts to the southeast, which connects Africa to Eurasia. It is the western part of the Fertile Crescent, the eastern part of the latter being Mesopotamia. This corridor is a land route of migrations of animals between Eurasia and Africa. In particular, it is believed that early hominins spread from Africa to Eurasia via the Levantine corridor and Horn of Africa. The corridor is named after the Levant.

Jebel Irhoud or Adrar n Ighoud (Standard Moroccan Tamazight: ⴰⴷⵔⴰⵔ ⵏ ⵉⵖⵓⴷ, romanized: Adrar n Iɣud; Arabic: جبل إيغود, Moroccan Arabic: žbəl iġud), is an archaeological site located just north of the town of Tlet Ighoud in Youssoufia Province, approximately 50 km (30 mi) south-east of the city of Safi in Morocco.

It is noted for the hominin fossils that have been found there since the discovery of the site in 1961. Originally thought to be Neanderthals, the specimens have since been assigned to Homo sapiens and, as reported in 2017, have been dated to roughly 300,000 years ago. This makes the remains by far the earliest known examples of Homo sapiens, and suggests that rather than arising in East Africa approximately 200,000 years ago, modern humans have been present across the length of Africa 100,000 years earlier.

The Omo remains are a collection of hominin bones discovered between 1967 and 1974 at the Omo Kibish sites near the Omo River, in Omo National Park in south-western Ethiopia. The bones were recovered by a scientific team from the Kenya National Museums directed by Richard Leakey and others. The remains from Kamoya's Hominid Site (KHS) were called Omo I and those from Paul I. Abell's Hominid Site (PHS) were called Omo II.

Australopithecus (/ˌɒstrələˈpɪθɪkəs, -loʊ-/, OS-trə-lə-PITH-i-kəs, -⁠loh-; or /ɒsˌtrələpɪˈθiːkəs/, os-TRA-lə-pi-THEE-kəs, from Latin austrālis 'southern' and Ancient Greek πίθηκος (píthēkos) 'ape') is a genus of early hominins that existed in Africa during the Pliocene and Early Pleistocene. The genera Homo (which includes modern humans), Paranthropus, and Kenyanthropus evolved from some Australopithecus species. Australopithecus is a member of the subtribe Australopithecina, which sometimes also includes Ardipithecus, though the term "australopithecine" is sometimes used to refer only to members of Australopithecus. Species include A. garhi, A. africanus, A. sediba, A. afarensis, A. anamensis, A. bahrelghazali, and A. deyiremeda. Debate exists as to whether some Australopithecus species should be reclassified into new genera, or if Paranthropus and Kenyanthropus are synonymous with Australopithecus, in part because of the taxonomic inconsistency.

Furthermore, because e.g. A. africanus is more closely related to humans, or their ancestors at the time, than e.g. A. anamensis and many more Australopithecus branches, Australopithecus cannot be consolidated into a coherent grouping without also including the genus Homo and other genera.

Paranthropus is a genus of extinct hominin which contains two widely accepted species: P. robustus and P. boisei. However, the validity of Paranthropus is contested, and it is sometimes considered to be synonymous with Australopithecus. They are also referred to as the robust australopithecines. They lived between approximately 2.9 and at least 1 million years ago (mya) from the end of the Pliocene to the Middle Pleistocene.

Paranthropus is characterised by robust skulls, with a prominent gorilla-like sagittal crest along the midline—which suggest strong chewing muscles—and broad, herbivorous teeth used for grinding. However, they likely preferred soft food over tough and hard food. Typically, Paranthropus species were generalist feeders, but while P. robustus was likely an omnivore, P. boisei seems to have been herbivorous, possibly preferring abundant bulbotubers. Paranthropoids were bipeds. Despite their robust heads, they had comparatively small bodies. Average weight and height are estimated to be 40 kg (88 lb) at 132 cm (4 ft) for P. robustus males, 50 kg (110 lb) at 137 cm (4 ft 6 in) for P. boisei males, 32 kg (71 lb) at 110 cm (3 ft 7 in) for P. robustus females, and 34 kg (75 lb) at 124 cm (4 ft 1 in) for P. boisei females.

Australopithecus africanus is an extinct species of australopithecine which lived between about 3.3 and 2.1 million years ago in the Late Pliocene to Early Pleistocene of South Africa. The species has been recovered from Taung, Sterkfontein, Makapansgat, and Gladysvale. The first specimen, the Taung child, was described by anatomist Raymond Dart in 1924, and was the first early hominin found. However, its closer relations to humans than to other apes would not become widely accepted until the middle of the century because most had believed humans evolved outside of Africa. It is unclear how A. africanus relates to other hominins, being variously placed as ancestral to Homo and Paranthropus, to just Paranthropus, or to just P. robustus. The specimen "Little Foot" is the most completely preserved early hominin, with 90% of the skeleton intact, and the oldest South African australopith. However, it is controversially suggested that it and similar specimens be split off into "A. prometheus".

A. africanus brain volume was about 420–510 cc (26–31 cu in). Like other early hominins, the cheek teeth were enlarged and had thick enamel. Male skulls may have been more robust than female skulls. Males may have been on average 140 cm (4 ft 7 in) in height and 40 kg (88 lb) in weight, and females 125 cm (4 ft 1 in) and 30 kg (66 lb). A. africanus was a competent biped, albeit less efficient at walking than humans. A. africanus also had several upper body traits in common with arboreal non-human apes. This is variously interpreted as either evidence of a partially or fully arboreal lifestyle, or as a non-functional vestige from a more apelike ancestor. The upper body of A. africanus is more apelike than that of the East African A. afarensis.

Homo habilis (lit. 'handy man') is an extinct species of archaic human from the Early Pleistocene of East and South Africa about 2.4 million years ago to 1.65 million years ago (mya). It is among the oldest species of archaic humans. Suggestions for pushing back the age to 2.8 Mya were made in 2015 based on the discovery of a jawbone. Upon species description in 1964, H. habilis was highly contested, with many researchers recommending it be synonymised with Australopithecus africanus, the only other early hominin known at the time, but H. habilis received more recognition as time went on and more relevant discoveries were made. By the 1980s, H. habilis was proposed to have been a human ancestor, directly evolving into Homo erectus, which directly led to modern humans. This viewpoint is now debated. Several specimens with insecure species identification were assigned to H. habilis, leading to arguments for splitting, namely into "H. rudolfensis" and "H. gautengensis" of which only the former has received wide support.

H. habilis brain size generally varied from 500 to 900 cm (31–55 cu in). The body proportions of H. habilis are only known from two highly fragmentary skeletons, and is based largely on assuming a similar anatomy to the earlier australopithecines. Because of this, it has also been proposed H. habilis be moved to the genus Australopithecus as Australopithecus habilis. However, the interpretation of H. habilis as a small-statured human with inefficient long-distance travel capabilities has been challenged. The presumed female specimen OH 62 is traditionally interpreted as having been 100–120 cm (3 ft 3 in – 3 ft 11 in) in height and 20–37 kg (44–82 lb) in weight assuming australopithecine-like proportions, but assuming humanlike proportions she would have been about 148 cm (4 ft 10 in) and 35 kg (77 lb). Nonetheless, Homo habilis may have been at least partially arboreal like what is postulated for australopithecines. Early hominins are typically reconstructed as having thick hair and marked sexual dimorphism with males much larger than females, though relative male and female size is not definitively known.

The Middle Stone Age (or MSA) was a period of African prehistory between the Early Stone Age and the Late Stone Age. It is generally considered to have begun around 280,000 years ago and ended around 50–25,000 years ago. The beginnings of particular MSA stone tools have their origins as far back as 550–500,000 years ago and as such some researchers consider this to be the beginnings of the MSA.

The MSA is often mistakenly understood to be synonymous with the Middle Paleolithic of Europe, especially due to their roughly contemporaneous time span; however, the Middle Paleolithic of Europe represents an entirely different hominin population, Homo neanderthalensis, than the MSA of Africa, which did not have Neanderthal populations. Additionally, current archaeological research in Africa has yielded much evidence to suggest that modern human behavior and cognition was beginning to develop much earlier in Africa during the MSA than it was in Europe during the Middle Paleolithic.

Prehistoric Arabia, or the prehistory of Arabia, refers to the era of the Arabian Peninsula that predates the start of recorded history in the region that came about at the dawn of Arabia's earliest kingdoms. Some define "Prehistoric Arabia" as the period between 10,000 and 1,000 BC, whereas others put its starting point all the way back to the time of the earliest hominin species two million years ago.

Research into Arabian prehistory has progressed slowly in comparison to that of other parts of the world, because of its harsh desert environment, historical restrictions in entry, and limited infrastructure for navigating the region, especially the desert interior. It has been said that the Arabian Peninsula "remains one of the last great unexplored regions of the ancient world".

Kenyanthropus ('man from Kenya') is a genus of extinct hominin identified from the Lomekwi site by Lake Turkana, Kenya, dated to 3.3 to 3.2 million years ago during the Middle Pliocene. It contains one species, K. platyops, but may also include the two-million-year-old Homo rudolfensis, or K. rudolfensis. Before its naming in 2001, Australopithecus afarensis was widely regarded as the only australopithecine to exist during the Middle Pliocene, but Kenyanthropus evinces a greater diversity than once acknowledged. Kenyanthropus is most recognisable by an unusually flat face and small teeth for such an early hominin, with values on the extremes or beyond the range of variation for australopithecines in regard to these features. Multiple australopithecine species may have coexisted by foraging for different food items (niche partitioning), which may be the reason why these apes anatomically differ in features related to chewing.

The Lomekwi site also yielded the earliest stone tool industry, the Lomekwian, characterised by the rudimentary production of simple flakes by pounding a core against an anvil or with a hammerstone. It may have been manufactured by Kenyanthropus, but it is unclear if multiple species were present at the site or not. The knappers were using volcanic rocks collected no more than 100 m (330 ft) from the site. Kenyanthropus seems to have lived on a lakeside or floodplain environment featuring forests and grasslands.

Ardipithecus is a genus of an extinct hominine that lived during the Late Miocene and Early Pliocene epochs in the Afar Depression, Ethiopia. Originally described as one of the earliest ancestors of humans after they diverged from the chimpanzees, the relation of this genus to human ancestors and whether it is a hominin is now a matter of debate. Two fossil species are described in the literature: A. ramidus, which lived about 4.4 million years ago during the early Pliocene, and A. kadabba, dated to approximately 5.6 million years ago (late Miocene). Initial behavioral analysis indicated that Ardipithecus could be very similar to chimpanzees; however, more recent analysis based on canine size and lack of canine sexual dimorphism indicates that Ardipithecus was characterised by reduced aggression, and that they more closely resemble bonobos.

Some analyses describe Australopithecus as being sister to Ardipithecus ramidus specifically. This means that Australopithecus is distinctly more closely related to Ardipithecus ramidus than Ardipithecus kadabba. Cladistically, then, Australopithecus (and eventually Homo sapiens) indeed emerged within the Ardipithecus lineage, and this lineage is not literally extinct.