History of life in the context of "Biological"

Biology is the scientific study of life and living organisms. It is a broad natural science that encompasses a wide range of fields and unifying principles that explain the structure, function, growth, origin, evolution, and distribution of life. Central to biology are five fundamental themes: the cell as the basic unit of life, genes and heredity as the basis of inheritance, evolution as the driver of biological diversity, energy transformation for sustaining life processes, and the maintenance of internal stability (homeostasis).

Biology examines life across multiple levels of organization, from molecules and cells to organisms, populations, and ecosystems. Subdisciplines include molecular biology, physiology, ecology, evolutionary biology, developmental biology, and systematics, among others. Each of these fields applies a range of methods to investigate biological phenomena, including observation, experimentation, and mathematical modeling. Modern biology is grounded in the theory of evolution by natural selection, first articulated by Charles Darwin, and in the molecular understanding of genes encoded in DNA. The discovery of the structure of DNA and advances in molecular genetics have transformed many areas of biology, leading to applications in medicine, agriculture, biotechnology, and environmental science.

The Phanerozoic is the current and the latest of the four geologic eons in the Earth's geologic time scale, covering the time period from 542 million years ago to the present. It is the eon during which abundant animal and plant life has proliferated, diversified and colonized various niches on the Earth's surface, beginning with the Cambrian period when animals first developed hard shells that can be clearly preserved in the fossil record. The time before the Phanerozoic, collectively called the Precambrian, is now divided into the Hadean, Archaean and Proterozoic eons.

The time span of the Phanerozoic starts with the sudden appearance of fossilised evidence of a number of animal phyla; the evolution of those phyla into diverse forms; the evolution of plants; the evolution of fish, arthropods and molluscs; the terrestrial colonization and evolution of insects, chelicerates, myriapods and tetrapods; and the development of modern flora dominated by vascular plants. During this time span, tectonic forces which move the continents had collected them into a single landmass known as Pangaea (the most recent supercontinent), which then separated into the current continental landmasses.

The Devonian (/dəˈvoʊni.ən, dɛ-/ də-VOH-nee-ən, deh-) is a geologic period and system of the Paleozoic era during the Phanerozoic eon, spanning 60.3 million years from the end of the preceding Silurian period at 419.62 million years ago (Ma), to the beginning of the succeeding Carboniferous period at 358.86 Ma. It is the fourth period of both the Paleozoic and the Phanerozoic. It is named after Devon, South West England, where rocks from this period were first studied.

The first significant evolutionary radiation of life on land occurred during the Devonian, as free-sporing land plants (pteridophytes) began to spread across dry land, forming extensive coal forests which covered the continents. By the middle of the Devonian, several groups of vascular plants had evolved leaves and true roots, and by the end of the period the first seed-bearing plants (pteridospermatophytes) appeared. This rapid evolution and colonization process, which had begun during the Silurian, is known as the Silurian-Devonian Terrestrial Revolution. The earliest land animals, predominantly arthropods such as myriapods, arachnids and hexapods, also became well-established early in this period, after beginning their colonization of land at least from the Ordovician Period.

Paleontology or palaeontology is the scientific study of the past, mainly but not exclusively through the study of fossils. Paleontologists use fossils as a means to classify organisms, measure geologic time, and assess the interactions between prehistoric organisms and their natural environment. While paleontological observations are known from at least the 6th century BC, the foundation of paleontology as a science dates back to the work of Georges Cuvier in 1796. Cuvier demonstrated evidence for the concept of extinction and how the life of the past was not necessarily the same as that of the present. The field developed rapidly over the course of the following decades, and the French word paléontologie was introduced for the study in 1822, which was derived from the Ancient Greek word for 'ancient' and words describing relatedness and a field of study. Further advances in the field accompanied the work of Charles Darwin who popularized the concept of evolution. Together, evolution and extinction can be understood as complementary processes that shaped the history of life.

Paleontology overlaps the most with the fields of geology and biology. It draws on technology and analysis of a wide range of sciences to apply them to the study of life and environments of the past, particularly for the subdisciplines of paleobiology and paleoecology that are analogous to biology and ecology. Paleontology also contributes to other sciences, being utilized for biostratigraphy to reconstruct the geologic time scale of Earth, or in studies on extinction to establish both external and internal factors that can lead to the disappearance of a species. Much of the history of life is now better understood because of advances in paleontology and the increase in interdisciplinary studies. Several improvements in understanding have occurred from the introduction of theoretical analysis to paleontology in the 1950s and 1960s which led to the rise of more focused fields of paleontology that assess the changing geography and climate of Earth, the phylogenetic relationships between different species, and the analysis of how fossilization occurs and what biases can impact the quality of the fossil record.

In biology, phylogenetics (/ˌfaɪloʊdʒəˈnɛtɪks, -lə-/) is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical data and observed heritable traits of DNA sequences, protein amino acid sequences, and morphology. The results are a phylogenetic tree—a diagram depicting the hypothetical relationships among the organisms, reflecting their inferred evolutionary history.

The tips of a phylogenetic tree represent the observed entities, which can be living taxa or fossils. A phylogenetic diagram can be rooted or unrooted. A rooted tree diagram indicates the hypothetical common ancestor of the taxa represented on the tree. An unrooted tree diagram (a network) makes no assumption about directionality of character state transformation, and does not show the origin or "root" of the taxa in question.

Background extinction rate (BER), also known as the normal extinction rate, refers to the standard rate of extinction in Earth's geological and biological history, excluding major extinction events, such as the current human-induced Holocene extinction. There have been five mass extinction events throughout Earth's history.