Harbin cranium in the context of "Harbin"

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⭐ Core Definition: Harbin cranium

The Harbin cranium is a nearly complete skull of an archaic human found in sediments of the Songhua River near Harbin on the Northeast China Plain, dating to at minimum 146,000 years ago during the Middle Pleistocene. It was described in 2021, and that year it was assigned as the holotype of the new human species Homo longi ("dragon man") named after Long Jiang, where the fossil skull was discovered . The Harbin cranium was initially hypothesized to belong to the same species as the Denisovans, and subsequent proteomics and mitochondrial DNA analyses confirmed its Denisovan affinities.

The Harbin cranium is broadly anatomically similar to other Middle Pleistocene Chinese specimens. Like other archaic humans, the skull is low and long, with massively developed brow ridges, wide eye sockets, and a large mouth. The skull is the longest ever found from any human species. Like modern humans and the much earlier Homo antecessor, the face is rather flat, but with a larger nose. The brain volume was 1,420 cc, within the range of modern humans and Neanderthals.

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Harbin cranium in the context of Denisovan

The Denisovans or Denisova hominins (/dəˈnsəvə/ də-NEE-sə-və) are an extinct species or subspecies of archaic human that ranged across Asia during the Middle to Late Pleistocene, approximately 200,000–32,000 years ago. Most of what is known about Denisovans comes from DNA evidence. While many recent fossils have been found and tentatively identified as Denisovan, the first Denisovans discovered were known from few physical remains. Consequently, no formal species name has been established. However, an analysis of the mitochondrial DNA and endogenous proteins from the Harbin cranium, which had been given the name Homo longi, showed with great certainty that this species represents a Denisovan.In a study published in September 2025, remains from six additional sites in China including the 1 million year old Yunxian man were proposed to be included in the species Homo longi along with the genetically confirmed Denisovans.

The first identification of a Denisovan individual occurred in 2010, based on mitochondrial DNA (mtDNA) extracted from a juvenile finger bone excavated from the Siberian Denisova Cave in the Altai Mountains in 2008. Nuclear DNA indicates close affinities with Neanderthals. The cave was also periodically inhabited by Neanderthals. Additional specimens from Denisova Cave were subsequently identified, as were specimens from the Baishiya Karst Cave on the Tibetan Plateau, Tam Ngu Hao 2 Cave in the Annamite Mountains of Laos, the Penghu channel between Taiwan and the mainland, and Harbin in Manchuria.

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