Haplodiploid in the context of Diploid

A wasp is any insect of the narrow-waisted suborder Apocrita of the order Hymenoptera which is neither a bee nor an ant; this excludes the broad-waisted sawflies (Symphyta), which look somewhat like wasps, but are in a separate suborder. The wasps do not constitute a clade, a complete natural group with a single ancestor, as bees and ants are deeply nested within the wasps, having evolved from wasp ancestors. Wasps that are members of the clade Aculeata can sting their prey.

The most commonly known wasps, such as yellowjackets and hornets, are in the family Vespidae and are eusocial, living together in a nest with an egg-laying queen and non-reproducing workers. Eusociality is favoured by the unusual haplodiploid system of sex determination in Hymenoptera, as it makes sisters exceptionally closely related to each other. However, the majority of wasp species are solitary, with each adult female living and breeding independently. Females typically have an ovipositor for laying eggs in or near a food source for the larvae, though in the Aculeata the ovipositor is often modified instead into a sting used for defense or prey capture.

Group selection is a proposed mechanism of evolution in which natural selection acts at the level of the group, instead of at the level of the individual or gene.

Early authors such as V. C. Wynne-Edwards and Konrad Lorenz argued that the behaviour of animals could affect their survival and reproduction as groups, speaking for instance of actions for the good of the species. In the 1930s, Ronald Fisher and J. B. S. Haldane proposed the concept of kin selection, a form of biological altruism from the gene-centered view of evolution, arguing that animals should sacrifice for their relatives, and thereby implying that they should not sacrifice for non-relatives. From the mid-1960s, evolutionary biologists such as John Maynard Smith, W. D. Hamilton, George C. Williams, and Richard Dawkins argued that natural selection acts primarily at the level of the gene. They argued on the basis of mathematical models that individuals would not altruistically sacrifice fitness for the sake of a group unless it would ultimately increase the likelihood of an individual passing on their genes. A consensus emerged that group selection did not occur, including in special situations such as the haplodiploid social insects like honeybees (in the Hymenoptera), where kin selection explains the behaviour of non-reproductives equally well, since the only way for them to reproduce their genes is via kin.