Fossil in the context of Biological species

The history of Europe is traditionally divided into four time periods: prehistoric Europe (prior to about 800 BC), classical antiquity (800 BC to AD 500), the Middle Ages (AD 500–1500), and the modern era (since AD 1500).

The first early European modern humans appear in the fossil record about 48,000 years ago, during the Paleolithic era. Settled agriculture marked the Neolithic era, which spread slowly across Europe from southeast to the north and west. The later Neolithic period saw the introduction of early metallurgy and the use of copper-based tools and weapons, and the building of megalithic structures, as exemplified by Stonehenge. During the Indo-European migrations, Europe saw migrations from the east and southeast. The period known as classical antiquity began with the emergence of the city-states of ancient Greece. Later, the Roman Empire came to dominate the entire Mediterranean Basin. The Migration Period of the Germanic people began in the late 4th century AD.

The Messinian salinity crisis (also referred to as the Messinian event, and in its latest stage as the Lago Mare event) was an event in which the Mediterranean Sea went into a cycle of partial or nearly complete desiccation (drying-up) throughout the latter part of the Messinian age of the Miocene epoch, from 5.96 to 5.33 Ma (million years ago). It ended with the Zanclean flood, when the Atlantic reclaimed the basin.

Sediment samples from below the deep seafloor of the Mediterranean Sea, which include evaporite minerals, soils, and fossil plants, show that the precursor of the Strait of Gibraltar closed about 5.96 million years ago, sealing the Mediterranean off from the Atlantic. This resulted in a period of partial desiccation of the Mediterranean Sea, the first of several such periods during the late Miocene. After the strait closed for the last time around 5.6 Ma, the region's generally dry climate at the time dried the Mediterranean basin out nearly completely within a thousand years. This massive desiccation left a deep dry basin, reaching 3 to 5 km (1.9 to 3.1 mi) deep below normal sea level, with a few hypersaline pockets similar to today's Dead Sea. Then, around 5.5 Ma, wetter climatic conditions resulted in the basin receiving more fresh water from rivers, progressively filling and diluting the hypersaline lakes into larger pockets of brackish water (much like today's Caspian Sea). The Messinian salinity crisis ended with the Strait of Gibraltar finally reopening 5.33 Ma, when the Atlantic rapidly filled up the Mediterranean basin in what is known as the Zanclean flood.

Archaeology or archeology is the study of human activity through the recovery and analysis of material culture. The archaeological record consists of artifacts, architecture, biofacts or ecofacts, sites, and cultural landscapes. Archaeology can be considered both a social science and a branch of the humanities. It is usually considered an independent academic discipline, but may also be classified as part of anthropology (in North America – the four-field approach), history or geography. The discipline involves surveying, excavation, and eventually analysis of data collected, to learn more about the past. In broad scope, archaeology relies on cross-disciplinary research.

Archaeologists study human prehistory and history, from the development of the first stone tools at Lomekwi in East Africa 3.3 million years ago up until recent decades. Archaeology is distinct from palaeontology, which is the study of fossil remains. Archaeology is particularly important for learning about prehistoric societies, for which, by definition, there are no written records. Prehistory includes over 99% of the human past, from the Paleolithic until the advent of literacy in societies around the world. Archaeology has various goals, which range from understanding culture history to reconstructing past lifeways to documenting and explaining changes in human societies through time. Derived from Greek, the term archaeology means "the study of ancient history".

A species (pl. species) is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. It can be defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. Other ways of defining species include their karyotype, DNA sequence, morphology, behaviour, or ecological niche. In addition, palaeontologists use the concept of the chronospecies since fossil reproduction cannot be examined. The most recent rigorous estimate for the total number of species of eukaryotes is between 8 and 8.7 million. About 14% of these had been described by 2011. All species (except viruses) are given a two-part name, a "binomen". The first part of a binomen is the name of a genus to which the species belongs. The second part is called the specific name or the specific epithet (in botanical nomenclature, also sometimes in zoological nomenclature). For example, Boa constrictor is one of the species of the genus Boa, with constrictor being the specific name.

While the definitions given above may seem adequate at first glance, when looked at more closely they represent problematic species concepts. For example, the boundaries between closely related species become unclear with hybridisation, in a species complex of hundreds of similar microspecies, and in a ring species. Also, among organisms that reproduce only asexually, the concept of a reproductive species breaks down, and each clonal lineage is potentially a microspecies. Although none of these are entirely satisfactory definitions, and while the concept of species may not be a perfect model of life, it is still a useful tool to scientists and conservationists for studying life on Earth, regardless of the theoretical difficulties. If species were fixed and distinct from one another, there would be no problem, but evolutionary processes cause species to change. This obliges taxonomists to decide, for example, when enough change has occurred to declare that a fossil lineage should be divided into multiple chronospecies, or when populations have diverged to have enough distinct character states to be described as cladistic species.

Genus (/ˈdʒiːnəs/; pl.: genera /ˈdʒɛnərə/) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses. In binomial nomenclature, the genus name forms the first part of the binomial species name for each species within the genus.

The composition of a genus is determined by taxonomists. The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including the idea that a newly defined genus should fulfill these three criteria to be descriptively useful:

Life is matter that has biological processes, such as signaling and the ability to sustain itself. It is defined descriptively by the capacity for homeostasis, organisation, metabolism, growth, adaptation, response to stimuli, and reproduction. All life over time eventually reaches a state of death, and none is immortal. Many philosophical definitions of living systems have been proposed, such as self-organizing systems. Defining life is further complicated by viruses, which replicate only in host cells, and the possibility of extraterrestrial life, which is likely to be very different from terrestrial life. Life exists all over the Earth in air, water, and soil, with many ecosystems forming the biosphere. Some of these are harsh environments occupied only by extremophiles. The life in a particular ecosystem is called its biota.

Life has been studied since ancient times, with theories such as Empedocles's materialism asserting that it was composed of four eternal elements, and Aristotle's hylomorphism asserting that living things have souls and embody both form and matter. Life originated at least 3.5 billion years ago, resulting in a universal common ancestor. This evolved into all the species that exist now, by way of many extinct species, some of which have left traces as fossils. Attempts to classify living things, too, began with Aristotle. Modern classification began with Carl Linnaeus's system of binomial nomenclature in the 1740s.

Limestone is a type of carbonate sedimentary rock which is the main source of the material lime. It is composed mostly of the minerals calcite and aragonite, which are different crystal forms of calcium carbonate CaCO₃. Limestone forms when these minerals precipitate out of water containing dissolved calcium. This can take place through both biological and nonbiological processes, though biological processes, such as the accumulation of corals and shells in the sea, have likely been more important for the last 540 million years. Limestone often contains fossils which provide scientists with information on ancient environments and on the evolution of life.

About 20% to 25% of sedimentary rock is carbonate rock, and most of this is limestone. The remaining carbonate rock is mostly dolomite, a closely related rock, which contains a high percentage of the mineral dolomite, CaMg(CO₃)₂. Magnesian limestone is an obsolete and poorly defined term used variously for dolomite, for limestone containing significant dolomite (dolomitic limestone), or for any other limestone containing a significant percentage of magnesium. Most limestone was formed in shallow marine environments, such as continental shelves or platforms, though smaller amounts were formed in many other environments. Much dolomite is secondary dolomite, formed by chemical alteration of limestone. Limestone is exposed over large regions of the Earth's surface, and because limestone is slightly soluble in rainwater, these exposures often are eroded to become karst landscapes. Most cave systems are found in limestone bedrock.

The Ediacaran (/ˌiːdiˈækərən/ EE-dee-ACK-ər-ən; formerly Vendian) biota is a taxonomic period classification that consists of all life forms that were present on Earth during the Ediacaran Period (c. 635–538.8 Mya). These were enigmatic tubular and frond-shaped, mostly sessile, organisms. Trace fossils of these organisms have been found worldwide, and represent the earliest known complex multicellular organisms. The term "Ediacara biota" has received criticism from some scientists due to its alleged inconsistency, arbitrary exclusion of certain fossils, and inability to be precisely defined.

The Ediacaran biota may have undergone evolutionary radiation in a proposed event called the Avalon explosion, 575 million years ago. This was after the Earth had thawed from the Cryogenian period's extensive glaciation. This biota largely disappeared with the rapid increase in biodiversity known as the Cambrian explosion. Most of the currently existing body plans of animals first appeared in the fossil record of the Cambrian rather than the Ediacaran. For macroorganisms, the Cambrian biota appears to have almost completely replaced the organisms that dominated the Ediacaran fossil record, although relationships are still a matter of debate.

The Ordovician (/ɔːrdəˈvɪʃi.ən, -doʊ-, -ˈvɪʃən/ or-də-VISH-ee-ən, -⁠doh-, -⁠VISH-ən) is a geologic period and system, the second of six periods of the Paleozoic Era, and the second of twelve periods of the Phanerozoic Eon. The Ordovician spans 41.6 million years from the end of the Cambrian Period 486.85 Ma (million years ago) to the start of the Silurian Period 443.1 Ma.

The Ordovician, named after the Welsh tribe of the Ordovices, was defined by Charles Lapworth in 1879 to resolve a dispute between followers of Adam Sedgwick and Roderick Murchison, who were placing the same rock beds in North Wales in the Cambrian and Silurian systems, respectively. Lapworth recognized that the fossil fauna in the disputed strata were different from those of either the Cambrian or the Silurian systems, and placed them in a system of their own. The Ordovician received international approval in 1960 (forty years after Lapworth's death), when it was adopted as an official period of the Paleozoic Era by the International Geological Congress.

Homo (from Latin homō 'human') is a genus of great ape (family Hominidae) that emerged from the early homininian genus Australopithecus, encompassing a single extant species, Homo sapiens (modern humans), along with a number of extinct species (e.g. Homo erectus and Homo neanderthalensis) classified as either ancestral or closely related to modern humans, collectively called archaic humans. Homo, together with the genus Paranthropus, is probably most closely related to the species Australopithecus africanus within Australopithecus. The closest living relatives of Homo are of the hominin genus Pan (chimpanzees and bonobos), with the ancestors of Pan and Homo estimated to have diverged around 5.7–11 million years ago during the Late Miocene.

The oldest member of the genus is Homo habilis, with fossil records of just over 2 million years ago. H. erectus appeared about 2 million years ago and spread throughout Africa (debatably as another species called Homo ergaster) and Eurasia in several migrations. The species was adaptive and successful, and persisted for more than a million years before gradually diverging into new species around 500,000 years ago.

Mollusca is a phylum of protostomic invertebrate animals, whose members are known as molluscs or mollusks (/ˈmɒləsks/). Around 76,000 extant species of molluscs are recognized, making it the second-largest animal phylum after Arthropoda. The number of additional fossil species is estimated between 60,000 and 100,000, and the proportion of undescribed species is very high. Many taxa remain poorly studied.

Molluscs are the largest marine phylum, comprising about 23% of all the named marine organisms. They are highly diverse, not just in size and anatomical structure, but also in behaviour and habitat, as numerous groups are freshwater and even terrestrial species. The phylum is typically divided into 7 or 8 taxonomic classes, of which two are entirely extinct. Cephalopod molluscs, such as squid, cuttlefish, and octopuses, are among the most neurologically advanced of all invertebrates—and either the giant squid or the colossal squid is the largest known extant invertebrate species. The gastropods (snails, slugs and abalone) are by far the most diverse class and account for 80% of the total classified molluscan species.

The geologic time scale or geological time scale (GTS) is a representation of time based on the rock record of Earth. It is a system of chronological dating that uses chronostratigraphy (the process of relating strata to time) and geochronology (a scientific branch of geology that aims to determine the age of rocks). It is used primarily by Earth scientists (including geologists, paleontologists, geophysicists, geochemists, and paleoclimatologists) to describe the timing and relationships of events in geologic history. The time scale has been developed through the study of rock layers and the observation of their relationships and identifying features such as lithologies, paleomagnetic properties, and fossils. The definition of standardised international units of geological time is the responsibility of the International Commission on Stratigraphy (ICS), a constituent body of the International Union of Geological Sciences (IUGS), whose primary objective is to precisely define global chronostratigraphic units of the International Chronostratigraphic Chart (ICC) that are used to define divisions of geological time. The chronostratigraphic divisions are in turn used to define geochronologic units.

Lithification (from the Ancient Greek word lithos meaning 'rock' and the Latin-derived suffix -ific) is the process in which sediments compact under pressure, expel connate fluids, and gradually become solid rock. Essentially, lithification is a process of porosity destruction through compaction and cementation. Lithification includes all the processes which convert unconsolidated sediments into sedimentary rocks. Petrifaction, though often used as a synonym, is more specifically used to describe the replacement of organic material by silica in the formation of fossils.

Pinnipeds (pronounced /ˈpɪnɪˌpɛdz/), commonly known as seals, are a widely distributed and diverse clade of carnivorous, fin-footed, semiaquatic, mostly marine mammals. They comprise the extant families Odobenidae (whose only living member is the walrus), Otariidae (the eared seals: sea lions and fur seals), and Phocidae (the earless seals, or true seals), with 34 extant species and more than 50 extinct species described from fossils. While seals were historically thought to have descended from two ancestral lines, molecular evidence supports them as a monophyletic group (descended from one ancestor). Pinnipeds belong to the suborder Caniformia of the order Carnivora; their closest living relatives are musteloids (weasels, raccoons, skunks and red pandas), having diverged about 50 million years ago.

Seals range in size from the 1 m (3 ft 3 in) and 45 kg (100 lb) Baikal seal to the 5 m (16 ft) and 3,200 kg (7,100 lb) southern elephant seal. Several species exhibit sexual dimorphism. They have streamlined bodies and four limbs that are modified into flippers. Though not as fast in the water as dolphins, seals are more flexible and agile. Otariids primarily use their front limbs to propel themselves through the water, while phocids and walruses primarily use their hind limbs for this purpose. Otariids and walruses have hind limbs that can be pulled under the body and used as legs on land. By comparison, terrestrial locomotion by phocids is more cumbersome. Otariids have visible external ears, while phocids and walruses lack these. Pinnipeds have well-developed senses—their eyesight and hearing are adapted for both air and water, and they have an advanced tactile system in their whiskers or vibrissae. Some species are well adapted for diving to great depths. They have a layer of fat, or blubber, under the skin to keep warm in cold water, and, other than the walrus, all species are covered in fur.

Jebel Irhoud or Adrar n Ighoud (Standard Moroccan Tamazight: ⴰⴷⵔⴰⵔ ⵏ ⵉⵖⵓⴷ, romanized: Adrar n Iɣud; Arabic: جبل إيغود, Moroccan Arabic: žbəl iġud), is an archaeological site located just north of the town of Tlet Ighoud in Youssoufia Province, approximately 50 km (30 mi) south-east of the city of Safi in Morocco.

It is noted for the hominin fossils that have been found there since the discovery of the site in 1961. Originally thought to be Neanderthals, the specimens have since been assigned to Homo sapiens and, as reported in 2017, have been dated to roughly 300,000 years ago. This makes the remains by far the earliest known examples of Homo sapiens, and suggests that rather than arising in East Africa approximately 200,000 years ago, modern humans have been present across the length of Africa 100,000 years earlier.