Flying and gliding animals in the context of "Amphibians"

Terrestrial animals are animals that live predominantly or entirely on land (e.g., cats, chickens, ants, most spiders), as compared with aquatic animals (e.g., fish, whales, octopuses, lobsters, etc.), who live predominantly or entirely in bodies of water; and semiaquatic animals (e.g., crocodilians, seals, platypus and most amphibians), who inhabit coastal, riparian or wetland areas and rely on both aquatic and terrestrial habitats. While most insects (who constitute over half of all known species in the animal kingdom) are terrestrial, some groups, such as mosquitoes and dragonflies, spend their egg and larval stages in water but emerge as fully terrestrial adults (imagos) after completing metamorphosis.

Terrestrial animals conduct respiratory gas exchange directly with the atmosphere, typically via specialized respiratory organs known as lungs, or via cutaneous respiration across the skin. They have also evolved homeostatic features such as impermeable cuticles that can restrict fluid loss, temperature fluctuations and infection, and an excretory system that can filter out nitrogenous waste in the form of urea or uric acid, in contrast to the ammonia-based excretion of aquatic animals. Without the buoyancy of an aqueous environment to support their weight, they have evolved robust skeletons that can hold up their body shape, as well as powerful appendages known as legs or limbs to facilitate terrestrial locomotion, although some perform limbless locomotion using body surface projections such as scales and setae. Some terrestrial animals even have wings or membranes that act as airfoils to generate lift, allowing them to fly and/or glide as airborne animals.

Ratites (/ˈrætaɪts/) are a polyphyletic group consisting of all birds within the infraclass Palaeognathae that lack keels and cannot fly. They are mostly large, long-necked, and long-legged, the exception being the kiwi, which is also the only nocturnal extant ratite.

The understanding of relationships within the paleognath clade has been in flux. Previously, all the flightless members had been assigned to the order Struthioniformes, which is more recently regarded as containing only the ostrich. The modern bird infraclass Palaeognathae consists of ratites and the flighted Neotropic tinamous (compare to Neognathae). Unlike other flightless birds, the ratites have no keel on their sternum—hence the name, from the Latin ratis ('raft', a vessel which has no keel—in contradistinction to extant flighted birds with a keel). Without this to anchor their wing muscles, they could not have flown even if they had developed suitable wings. Ratites are a polyphyletic group; tinamous fall within them, and are the sister group of the extinct moa. This implies that flightlessness is a trait that evolved independently multiple times in different ratite lineages.

Pterosaurs are an extinct clade of flying reptiles in the order Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 million to 66 million years ago). Pterosaurs are the earliest vertebrates known to have evolved powered flight. Their wings were formed by a membrane of skin, muscle, and other tissues stretching from the ankles to a dramatically lengthened fourth finger.

Traditionally, pterosaurs were divided into two major types. Basal pterosaurs (also called non-pterodactyloid pterosaurs or 'rhamphorhynchoids') were smaller animals, up to two meter wingspan, with fully toothed jaws and, typically, long tails. Their wide wing membranes probably included and connected the hindlimbs. On the ground, they would have had an awkward sprawling posture due to short metacarpals, but the anatomy of their joints and strong claws would have made them effective climbers, and some may have lived in trees. Basal pterosaurs were insectivores, piscivores or predators of small land vertebrates. Later pterosaurs (pterodactyloids) evolved many sizes, shapes, and lifestyles. Pterodactyloids had narrower wings with free hindlimbs, highly reduced tails, and long necks with large heads. On the ground, they walked well on all four limbs due to long metacarpals with an upright posture, standing plantigrade on the hind feet and folding the wing finger upward to walk on the metacarpals with the three smaller fingers of the hand pointing to the rear. They could take off from the ground, and fossil trackways show that at least some species were able to run, wade, and/or swim. Their jaws had horny beaks, and some groups lacked teeth. Some groups developed elaborate head crests with sexual dimorphism. Since 2010 it is understood that many species, the basal Monofenestrata, were intermediate in build, combining an advanced long skull with long tails.

Flight or flying is the motion of an object through an atmosphere or through the vacuum of space, in this case also called spaceflight, without contacting any planetary surface. This can be achieved by generating aerodynamic lift associated with gliding or propulsive thrust, aerostatically using buoyancy, or by ballistic movement.

Many things can fly, from animal aviators such as birds, bats and insects, to natural gliders/parachuters such as patagial animals, anemochorous seeds and ballistospores, to human inventions like aircraft (airplanes, helicopters, airships, balloons, etc.) and rockets which may propel spacecraft and spaceplanes.

The patagium (pl.: patagia) is a membranous body part that assists an animal in obtaining lift when gliding or flying. The structure is found in extant and extinct groups of flying and gliding animals including bats, theropod dinosaurs (including birds and some dromaeosaurs), pterosaurs, gliding mammals, some flying lizards, and flying frogs. The patagium that stretches between an animal's hind limbs is called the uropatagium (especially in bats) or the interfemoral membrane.

Gliding flight is heavier-than-air flight without the use of thrust; the term volplaning also refers to this mode of flight in animals. It is employed by gliding animals and by aircraft such as gliders. This mode of flight involves flying a significant distance horizontally compared to its descent and therefore can be distinguished from a mostly straight downward descent like a round parachute.

Although the human application of gliding flight usually refers to aircraft designed for this purpose, most powered aircraft are capable of gliding without engine power. As with sustained flight, gliding generally requires the application of an airfoil, such as the wings on aircraft or birds, or the gliding membrane of a gliding possum. However, gliding can be achieved with a flat (uncambered) wing, as with a simple paper plane, or even with card-throwing. However, some aircraft with lifting bodies and animals such as the flying snake can achieve gliding flight without any wings by creating a flattened surface underneath.