Flightless bird in the context of "Burrowing owl"

Birds are a group of warm-blooded theropod dinosaurs constituting the class Aves, characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweight skeleton. Birds live worldwide and range in size from the 5.5 cm (2.2 in) bee hummingbird to the 2.8 m (9 ft 2 in) common ostrich. There are over 11,000 living species and they are split into 44 orders. More than half are passerine or "perching" birds. Birds have wings whose development varies according to species; the only known groups without wings are the extinct moa and elephant birds. Wings, which are modified forelimbs, gave birds the ability to fly, although further evolution has led to the loss of flight in some birds, including ratites, penguins, and diverse endemic island species. The digestive and respiratory systems of birds are also uniquely adapted for flight. Some bird species of aquatic environments, particularly seabirds and some waterbirds, have further evolved for swimming. The study of birds is called ornithology.

Birds evolved from earlier theropods, and thus constitute the only known living dinosaurs. Likewise, birds are considered reptiles in the modern cladistic sense of the term, and their closest living relatives are the crocodilians. Birds are descendants of the primitive avialans (whose members include Archaeopteryx) which first appeared during the Late Jurassic. According to some estimates, modern birds (Neornithes) evolved in the Late Cretaceous or between the Early and Late Cretaceous (100 Ma) and diversified dramatically around the time of the Cretaceous–Paleogene extinction event 66 million years ago, which killed off the pterosaurs and all non-ornithuran dinosaurs.

Penguins are a group of flightless, semi-aquatic, sea birds which live almost exclusively in the Southern Hemisphere. Only one species, the Galápagos penguin, lives at, and slightly north of, the equator. Highly adapted for life in the ocean water, penguins have countershaded dark and white plumage and flippers for swimming. Most penguins feed on krill, fish, squid and other forms of sea life which they catch with their bills and swallow whole while swimming. A penguin has a spiny tongue and powerful jaws to grip slippery prey.

They spend about half of their lives on land and the other half in the sea. The largest living species is the emperor penguin (Aptenodytes forsteri): on average, adults are about 1.1 m (3 ft 7 in) tall and weigh 35 kg (77 lb). The smallest penguin species is the little blue penguin (Eudyptula minor), also known as the fairy penguin, which stands around 30–33 cm (12–13 in) tall and weighs 1.2–1.3 kg (2.6–2.9 lb). Today, larger penguins generally inhabit colder regions, and smaller penguins inhabit regions with temperate or tropical climates. Some prehistoric penguin species were enormous: as tall or heavy as an adult human. There was a great diversity of species in subantarctic regions, and at least one giant species in a region around 2,000 km south of the equator 35 mya, during the Late Eocene, a climate decidedly warmer than today.

Ostriches are large flightless birds. Two living species are recognised; the common ostrich, native to large parts of Sub-Saharan Africa, and the Somali ostrich, native to the Horn of Africa.

They are the heaviest and largest living birds, with adult common ostriches weighing anywhere between 63.5 and 145 kilograms and laying the largest eggs of any living land animal. With the ability to run at 70 km/h (43.5 mph), they are the fastest birds on land. They are farmed worldwide, with significant industries in the Philippines and in Namibia. South Africa produces about 70% of global ostrich products, with the industry largely centered around the town of Oudtshoorn. Ostrich leather is a lucrative commodity, and the large feathers are used as plumes for the decoration of ceremonial headgear. Ostrich eggs and meat have been used by humans for millennia. Ostrich oil is another product that is made using ostrich fat.

The common ostrich (Struthio camelus), or simply ostrich, is a species of flightless bird native to certain areas of Africa. It is one of two extant species of ostriches, the only living members of the genus Struthio in the ratite group of birds. The other is the Somali ostrich (Struthio molybdophanes), which has been recognized as a distinct species by BirdLife International since 2014, having been previously considered a distinctive subspecies of ostrich.

The common ostrich belongs to the order Struthioniformes. Struthioniformes previously contained all the ratites, such as the kiwis, emus, rheas, and cassowaries. However, recent genetic analysis has found that the group is not monophyletic, as it is paraphyletic with respect to the tinamous, so the ostriches are now classified as the only members of the order. Phylogenetic studies have shown that it is the sister group to all other members of Palaeognathae, and thus the flighted tinamous are the sister group to the extinct moa. It is distinctive in its appearance, with a long neck and legs, and can run for a long time at a speed of 55 km/h (34 mph) with short bursts up to about 70 km/h (43 mph), the fastest land speed of any bipedal animal. The common ostrich is the largest living species of bird and thus the largest living dinosaur. It lays the largest eggs of any living bird (the extinct giant elephant bird (Aepyornis maximus) of Madagascar and the south island giant moa (Dinornis robustus) of New Zealand laid larger eggs).

Moa (order Dinornithiformes) are an extinct group of flightless birds formerly endemic to New Zealand. During the Late Pleistocene-Holocene, there were nine species (in six genera). The two largest species, Dinornis robustus and Dinornis novaezelandiae, reached about 3.6 metres (12 ft) in height with neck outstretched, and weighed about 230 kilograms (510 lb) while the smallest, the bush moa (Anomalopteryx didiformis), was around the size of a turkey. Estimates of the moa population when Polynesians settled New Zealand circa 1300 vary between 58,000 and approximately 2.5 million.

Moa are traditionally placed in the ratite group. Genetic studies have found that their closest relatives are the flighted South American tinamous, once considered a sister group to ratites. The nine species of moa were the only wingless birds, lacking even the vestigial wings that all other ratites have. They were the largest terrestrial animals and dominant herbivores in New Zealand's forest, shrubland, and subalpine ecosystems until the arrival of the Māori, and were hunted only by Haast's eagle. Moa extinction occurred within 100 years of human settlement of New Zealand, primarily due to overhunting.

Ratites (/ˈrætaɪts/) are a polyphyletic group consisting of all birds within the infraclass Palaeognathae that lack keels and cannot fly. They are mostly large, long-necked, and long-legged, the exception being the kiwi, which is also the only nocturnal extant ratite.

The understanding of relationships within the paleognath clade has been in flux. Previously, all the flightless members had been assigned to the order Struthioniformes, which is more recently regarded as containing only the ostrich. The modern bird infraclass Palaeognathae consists of ratites and the flighted Neotropic tinamous (compare to Neognathae). Unlike other flightless birds, the ratites have no keel on their sternum—hence the name, from the Latin ratis ('raft', a vessel which has no keel—in contradistinction to extant flighted birds with a keel). Without this to anchor their wing muscles, they could not have flown even if they had developed suitable wings. Ratites are a polyphyletic group; tinamous fall within them, and are the sister group of the extinct moa. This implies that flightlessness is a trait that evolved independently multiple times in different ratite lineages.

The Great American Biotic Interchange (commonly abbreviated as GABI), also known as the Great American Interchange and the Great American Faunal Interchange, was an important late Cenozoic paleozoogeographic biotic interchange event in which land and freshwater fauna migrated from North America to South America via Central America and vice versa, as the volcanic Isthmus of Panama rose up from the sea floor, forming a land bridge between the previously separated continents. Although earlier dispersals had occurred, probably over water, the migration accelerated dramatically about 2.7 million years (Ma) ago during the Piacenzian age. It resulted from the joining of the Neotropic (roughly South American) and Nearctic (roughly North American) biogeographic realms definitively to form the Americas. The interchange is visible from observation of both biostratigraphy and nature (neontology). Its most dramatic effect is on the zoogeography of mammals, but it also gave an opportunity for reptiles, amphibians, arthropods, weak-flying or flightless birds, and even freshwater fish to migrate. Coastal and marine biota were affected in the opposite manner; the formation of the Central American Isthmus caused what has been termed the Great American Schism, with significant diversification and extinction occurring as a result of the isolation of the Caribbean from the Pacific.

The occurrence of the interchange was first discussed in 1876 by the "father of biogeography", Alfred Russel Wallace. Wallace had spent five years exploring and collecting specimens in the Amazon basin. Others who made significant contributions to understanding the event in the century that followed include Florentino Ameghino, W. D. Matthew, W. B. Scott, Bryan Patterson, George Gaylord Simpson and S. David Webb. The Pliocene timing of the formation of the connection between North and South America was discussed in 1910 by Henry Fairfield Osborn.

The dodo (Raphus cucullatus) is an extinct flightless bird that was endemic to Mauritius, an island east of Madagascar in the Indian Ocean. The dodo's closest relative was the also-extinct and flightless Rodrigues solitaire. The two formed the subtribe Raphina, a clade of extinct flightless birds that are a part of the group that includes pigeons and doves (the family Columbidae). The closest living relative of the dodo is the Nicobar pigeon. A white dodo was once thought to have existed on the nearby island of Réunion, but it is now believed that this assumption was merely confusion based on the also-extinct Réunion ibis and paintings of white dodos.

Subfossil remains show the dodo measured about 62.6–75 centimetres (2.05–2.46 ft) in height and may have weighed 10.6–17.5 kg (23–39 lb) in the wild. The dodo's appearance in life is evidenced only by drawings, paintings, and written accounts from the 17th century. Since these portraits vary considerably, and since only some of the illustrations are known to have been drawn from live specimens, the dodos' exact appearance in life remains unresolved, and little is known about its behaviour. It has been depicted with brownish-grey plumage, yellow feet, a tuft of tail feathers, a grey, naked head, and a black, yellow, and green beak. It used gizzard stones to help digest its food, which is thought to have included fruits, and its main habitat is believed to have been the woods in the drier coastal areas of Mauritius. One account states its clutch consisted of a single egg. It is presumed that the dodo became flightless because of the ready availability of abundant food sources and a relative absence of predators on Mauritius. Though the dodo has historically been portrayed as being fat and clumsy, it is now thought to have been well-adapted for its ecosystem.