Ephedra (plant) in the context of "Temperate climates"

A vessel element or vessel member (also called a xylem vessel) is one of the cell types found in xylem, the water conducting tissue of plants. Vessel elements are found in most angiosperms (flowering plants) and in some gymnosperms such as cycads and Ephedra, but absent in conifers. Vessel elements are the main feature distinguishing the "hardwood" of angiosperms from the "softwood" of conifers.

Gnetophyta (/nɛˈtɒfɪtə, ˈnɛtoʊfaɪtə/) is a division of plants (alternatively considered the subclass Gnetidae or order Gnetales), grouped within the gymnosperms (which also includes conifers, cycads, and ginkgos), that consists of some 70 species across the three relict genera: Gnetum (family Gnetaceae), Welwitschia (family Welwitschiaceae), and Ephedra (family Ephedraceae). The earliest unambiguous records of the group date to the Jurassic, and they achieved their highest diversity during the Early Cretaceous. The primary difference between gnetophytes and other gymnosperms is the presence of vessel elements, a system of small tubes (xylem) that transport water within the plant, similar to those found in flowering plants. Because of this, gnetophytes were once thought to be the closest gymnosperm relatives to flowering plants, but more recent molecular studies have brought this hypothesis into question, with many recent phylogenies finding them to be nested within the conifers.

Though it is clear they are all related, the exact evolutionary inter-relationships between gnetophytes are unclear. Some classifications hold that all three genera should be placed in a single order (Gnetales), while other classifications say they should be distributed among three separate orders, each containing a single family and genus. Most morphological and molecular studies confirm that the genera Gnetum and Welwitschia diverged from each other more recently than they did from Ephedra.

Gnetum is a genus of gymnosperms, the sole genus in the family Gnetaceae within the Gnetophyta. They are tropical evergreen trees, shrubs and lianas. Unlike other gymnosperms, they possess vessel elements in the xylem. Some species have been proposed to have been the first plants to be insect-pollinated as their fossils occur in association with extinct pollinating scorpionflies. Molecular phylogenies based on nuclear and plastid sequences from most of the species indicate hybridization among some of the Southeast Asian species. Fossil-calibrated molecular-clocks suggest that the Gnetum lineages now found in Africa, South America and Southeast Asia are the result of ancient long-distance dispersal across seawater.

Their leaves are rich in phytochemicals such as flavonoids and stilbenes. Of the species studied so far, Gnetum have photosynthetic and transpiration capacities which are considerably lower than those of other seed plants, due to the absence of multiple chloroplast genes essential for photosynthesis, a trait they seem to share with the other living members of Gnetophyta, Ephedra and Welwitschia, as well as conifers. There are over 50 different species of Gnetum.

Welwitschia is a monotypic genus of gnetophytes containing only the species Welwitschia mirabilis. It is named after the Austrian botanist Friedrich Welwitsch, who documented the plant in the 1850s. In common use, it is sometimes referred to as the tree tumbo. It is native to Angola and Namibia, where it grows in the extreme conditions of the Namib desert, tolerating high heat and low precipitation. Welwitschia is the only living genus of the family Welwitschiaceae and order Welwitschiales, and is one of three extant genera of gnetophytes, alongside Gnetum and Ephedra.

Welwitschia is well known for its unique morphology. The plant has only two leaves that grow out of a large woody stem, known as a crown, and continue to grow throughout the plant's life. Welwitschia is one of the longest-living plants on Earth, with some individuals being thousands of years old. Because of the long lifespan the leaves can become quite large, often reaching several meters in length.

Ephedraceae is a family of gymnosperms belonging to Gnetophyta, it contains only a single extant genus, Ephedra, as well as a number of extinct genera from the Early Cretaceous.