Endosymbiosis in the context of "Endosymbiotic theory"

An aquatic animal is any animal, whether vertebrate or invertebrate, that lives in a body of water for all or most of its lifetime. Aquatic animals generally conduct aquatic respiration by extracting dissolved oxygen in water via specialised respiratory organs called gills, through the skin or across enteral mucosae, although some are secondarily aquatic animals (e.g. marine reptiles and marine mammals) evolved from terrestrial ancestors that re-adapted to aquatic environments, in which case they actually use lungs to breathe air and are essentially holding their breath when living in water. Some species of gastropod mollusc, such as the eastern emerald sea slug, are even capable of kleptoplastic photosynthesis via endosymbiosis with ingested yellow-green algae.

Almost all aquatic animals reproduce in water, either oviparously or viviparously, and many species routinely migrate between different water bodies during their life cycle. Some animals have fully aquatic life stages (typically as eggs and larvae), while as adults they become terrestrial or semi-aquatic after undergoing metamorphosis. Such examples include amphibians such as frogs, many flying insects such as mosquitoes, mayflies, dragonflies, damselflies and caddisflies, as well as some species of cephalopod molluscs such as the algae octopus (whose larvae are completely planktonic, but adults are highly terrestrial).

Plants are the eukaryotes that comprise the kingdom Plantae; they are predominantly photosynthetic. This means that they obtain their energy from sunlight, using chloroplasts derived from endosymbiosis with cyanobacteria to produce sugars from carbon dioxide and water, using the green pigment chlorophyll. Exceptions are parasitic plants that have lost the genes for chlorophyll and photosynthesis, and obtain their energy from other plants or fungi. Most plants are multicellular, except for some green algae.

Historically, as in Aristotle's biology, the plant kingdom encompassed all living things that were not animals, and included algae and fungi. Definitions have narrowed since then; current definitions exclude fungi and some of the algae. By the definition used in this article, plants form the clade Viridiplantae (green plants), which consists of the green algae and the embryophytes or land plants (hornworts, liverworts, mosses, lycophytes, ferns, conifers and other gymnosperms, and flowering plants). A definition based on genomes includes the Viridiplantae, along with the red algae and the glaucophytes, in the clade Archaeplastida.

Viridiplantae (lit. 'green plants'; kingdom Plantae sensu stricto) is a clade of around 450,000–500,000 species of eukaryotic organisms, most of which obtain their energy by photosynthesis. The green plants are chloroplast-bearing autotrophs that play important primary production roles in both terrestrial and aquatic ecosystems. They include green algae, which are primarily aquatic, and the land plants (embryophytes, Plantae sensu strictissimo), which emerged within freshwater green algae. Green algae traditionally excludes the land plants, rendering them a paraphyletic group, however it is cladistically accurate to think of land plants as a special clade of green algae that evolved to thrive on dry land. Since the realization that the embryophytes emerged from within the green algae, some authors are starting to include them.

Viridiplantae species all have cells with cellulose in their cell walls, and primary chloroplasts derived from endosymbiosis with cyanobacteria that contain chlorophylls a and b and lack phycobilins. Corroborating this, a basal phagotroph Archaeplastida group has been found in the Rhodelphidia. In some classification systems, the group has been treated as a kingdom, under various names, e.g. Viridiplantae, Chlorobionta, or simply Plantae, the latter expanding the traditional plant kingdom of embryophytes to include the green algae. Adl et al., who produced a classification for all eukaryotes in 2005, introduced the name Chloroplastida for this group, reflecting the group having primary chloroplasts. They rejected the name Viridiplantae on the grounds that some of the species are not plants as understood traditionally. Together with Rhodophyta, glaucophytes and other basal groups, Viridiplantae belong to a larger clade called Archaeplastida which in itself is sometimes described as Plantae sensu lato.

The evolution of plants has resulted in a wide range of complexity, from the earliest algal mats of unicellular archaeplastids evolved through endosymbiosis, through multicellular marine and freshwater green algae, to spore-bearing terrestrial bryophytes, lycopods and ferns, and eventually to the complex seed-bearing gymnosperms and angiosperms (flowering plants) of today. While many of the earliest groups continue to thrive, as exemplified by red and green algae in marine environments, more recently derived groups have displaced previously ecologically dominant ones; for example, the ascendance of flowering plants over gymnosperms in terrestrial environments.

There is evidence that cyanobacteria and multicellular thalloid eukaryotes lived in freshwater communities on land as early as 1 billion years ago, and that communities of complex, multicellular photosynthesizing organisms existed on land in the late Precambrian, around 850 million years ago.

Euglenids or euglenoids are one of the best-known groups of eukaryotic flagellates: single-celled organisms with flagella, or whip-like tails. They are classified in the phylum Euglenozoa, class Euglenida or Euglenoidea. Euglenids are commonly found in fresh water, especially when it is rich in organic materials, but they have a few marine and endosymbiotic members. Many euglenids feed by phagocytosis, or strictly by diffusion. A monophyletic subgroup known as Euglenophyceae have chloroplasts and produce their own food through photosynthesis. This group contains the carbohydrate paramylon.

Euglenids split from other Euglenozoa (a larger group of flagellates) more than a billion years ago. The plastids (membranous organelles) in all extant photosynthetic species result from secondary endosymbiosis between a euglenid and a green alga.

Chlorella vulgaris is a species of green microalga in the division Chlorophyta. This unicellular alga was discovered in 1890 by Martinus Willem Beijerinck as the first microalga with a well-defined nucleus. It is the type species of the genus Chlorella. It is found in freshwater and terrestrial habitats, and has a cosmopolitan distribution.

Chlorella vulgaris has a number of potential applications in science, such as biofuel, livestock feed, and wastewater treatment. Beginning in the 1990s, German scientists noticed the high protein content of C. vulgaris and began to consider it as a new food source. Japan is currently the largest consumer of Chlorella, both for nutritional and therapeutic purposes, and it is used as a dietary supplement or protein-rich food additive in several countries worldwide.