Dorsal column–medial lemniscus pathway in the context of "Dorsal column nuclei"

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Dorsal column–medial lemniscus pathway in the context of Brook trout

The brook trout (Salvelinus fontinalis) is a species of freshwater fish in the char genus Salvelinus of the salmon family Salmonidae native to Eastern North America in the United States and Canada. Two ecological forms of brook trout have been recognized by the US Forest Service. One ecological form is long-lived potamodromous populations in Lake Superior known as coaster trout or coasters. The second ecological form is the short-living predaceous anadromous populations which are found in northern lakes and coastal rivers from Long Island to Hudson Bay, which are referred to as salters. In parts of its range, it is also known as the eastern brook trout, speckled trout, brook char (or charr), squaretail, brookie, or mud trout, among others. Adult coaster brook trout are capable of reaching sizes over 2 feet in length and weigh up to 6.8 kg (15 lb), whereas adult salters average between 6 and 15 inches in length and weigh between 0.5 and 2.3 kg (1 and 5 lb). The brook trout is characterized by its distinctive olive-green body with yellow and blue-rimmed red spots, white and black edged orange fins, and dorsal vermiculation. The diet of the brook trout is restrictive to the season and location of the fish, but will typically consist of terrestrial and aquatic insects, fry, crustaceans, zooplankton, and worms.

Throughout history, non-native brook trout have been transplanted beyond its native borders, where it has spread across North America and much of the world. These brook trout have been introduced since the 1800s by means of artificial propagation and aquaculture in hope of promoting fishery resources. Through this transplantation, brook trout have been observed to affect native populations by outcompeting, preying upon, and hybridizing with many native aquatic species. This invasive nature via human-mediated introductory has led to their classification in the list of the top 100 globally invasive species.

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Dorsal column–medial lemniscus pathway in the context of Dorsal columns

The dorsal column nuclei are a pair of nuclei in the dorsal columns of the dorsal column–medial lemniscus pathway (DCML) in the brainstem. The name refers collectively to the cuneate nucleus and gracile nucleus, which are situated at the lower end of the medulla oblongata. Both nuclei contain second-order neurons of the DCML, which convey fine touch and proprioceptive information from the body to the brain via the thalamus.

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Dorsal column–medial lemniscus pathway in the context of Sensory decussation

The sensory decussation or decussation of the lemnisci is a decussation (a crossing over) of axons from the gracile nucleus and cuneate nucleus, known together as the dorsal column nuclei. The dorsal column nuclei are responsible for fine touch, vibration, proprioception and two-point discrimination.

The fibers of this decussation are called the internal arcuate fibers and are found at the superior aspect of the closed medulla oblongata, superior to the motor decussation. Neurons of these nuclei are second-order neurons in the dorsal column–medial lemniscus pathway.

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Dorsal column–medial lemniscus pathway in the context of Medial lemniscus

The medial lemniscus, also known as Reil's band or Reil's ribbon (for German anatomist Johann Christian Reil), is a large ascending bundle of heavily myelinated axons that decussate in the brainstem, specifically in the medulla oblongata. The medial lemniscus is formed by the crossings of the internal arcuate fibers. The internal arcuate fibers are composed of axons of the gracile nucleus and the cuneate nucleus. The cell bodies of the nuclei lie contralaterally.

The medial lemniscus is part of the somatosensory dorsal column–medial lemniscus pathway, which ascends in the spinal cord to the thalamus. Lesions of the medial lemniscus cause an impairment of vibratory and touch-pressure sense.

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