Compsognathidae in the context of Juravenator


Compsognathidae in the context of Juravenator

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⭐ Core Definition: Compsognathidae

Compsognathidae is a potentially polyphyletic family of coelurosaurian theropod dinosaurs. Compsognathids were small carnivores, generally conservative in form, hailing from the Jurassic and Cretaceous Periods. The bird-like features of these species, along with other dinosaurs such as Archaeopteryx inspired the idea for the connection between dinosaur reptiles and modern-day avian species. Compsognathid fossils preserve diverse integument — skin impressions are known from four genera commonly placed in the group, Compsognathus, Sinosauropteryx, Sinocalliopteryx and Juravenator. While the latter three show evidence of a covering of some of the earliest primitive feathers over much of the body, Juravenator and Compsognathus also show evidence of scales on the tail or hind legs. "Ubirajara jubatus", informally described in 2020, had elaborate integumentary structures on its back and shoulders superficially similar to the display feathers of a standardwing bird-of-paradise, and unlike any other non-avian dinosaur currently described.

The first member of the group, Compsognathus, was discovered in 1861, after Johann A. Wagner published his description of the taxon. The family was created by Edward Drinker Cope in 1875. This classification was accepted by Othniel Charles Marsh in 1882, and added to the Coelurosauria clade by Friedrich von Huene in 1914 after additional fossils had been found. With further discoveries, fossils have been uncovered across three different continents, in Asia, Europe, and South America. Assignment to Compsognathidae is usually determined through examination of the metacarpal, which is used to separate Compsognathidae from other dinosaurs. However, classification is still complicated due to similarities to the body of several other theropod dinosaurs, as well as the lack of unifying, diagnostic features that are shared by all compsognathids. Some authors have proposed that Compsognathidae is not a monophyletic group, and at least some compsognathids represent juvenile specimens of larger tetanuran theropods, such as carnosaurs and tyrannosaurs.

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Compsognathidae in the context of Coelurosauria

Coelurosauria (/sɪˌljʊərəˈsɔːri.ə/; from Greek, meaning "hollow-tailed lizards") is the clade containing all theropod dinosaurs more closely related to birds than to carnosaurs.

Coelurosauria is a subgroup of theropod dinosaurs that includes compsognathids, tyrannosauroids, ornithomimosaurs, maniraptorans, and over the recent years, megaraptorans (although their position within the clade is unclear). Maniraptora includes birds, the only known dinosaur group alive today. In the past, Coelurosauria was used to refer to all small theropods, but this classification has since been amended.

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Compsognathidae in the context of Megaraptora

Megaraptora is a clade of carnivorous theropod dinosaurs. Its derived members, the Megaraptoridae are noted for their large hand claws and powerfully-built forelimbs, which are usually reduced in size in other large theropods. Although undoubtedly members of the clade Tetanurae, their relationships to others members of this group have been subject to dispute.

Megaraptorans are incompletely known, and no complete megaraptoran skeleton has been found. However, they still possessed a number of unique features. Their forelimbs were large and strongly built, and the ulna bone had a unique shape in members of the family Megaraptoridae, a subset of megaraptorans which excludes Fukuiraptor and Phuwiangvenator. The first two fingers were elongated, with massive curved claws, while the third finger was small. Megaraptoran skull material is very incomplete, but a juvenile Megaraptor described in 2014 preserved a portion of the snout, which was long and slender. Leg bones referred to megaraptorans were also quite slender and similar to those of coelurosaurs adapted for running. Although megaraptorans were thick-bodied theropods, their bones were heavily pneumatized, or filled with air pockets. The vertebrae, ribs, and the ilium bone of the hip were pneumatized to an extent which was very rare among theropods, only seen elsewhere in taxa such as Neovenator. Other characteristic features include opisthocoelous neck vertebrae and compsognathid-like teeth.

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