Chirotherium, also known as Cheirotherium (‘hand-beast’), is a Triassic trace fossil consisting of five-fingered (pentadactyle) footprints and whole tracks. These look, by coincidence, remarkably like the hands of apes and bears, with the outermost toe having evolved to extend out to the side like a thumb, although probably only functioning to provide a firmer grip in mud. Chirotherium tracks were first found in 1834 in Lower Triassic sandstone (Buntsandstein) in Thuringia, Germany, dating from about 243 million years ago (mya).
The creatures who made the footprints and tracks were probably pseudosuchian archosaurs related to the ancestors of the crocodiles. They likely belonged to either prestosuchidae or rauisuchidae groups, which were both large carnivores with semi-erect gaits.
