Arachnids are arthropods in the class Arachnida (/əˈræknɪdə/) of the subphylum Chelicerata. Arachnida includes, among others, spiders, scorpions, ticks, mites, pseudoscorpions, harvestmen, camel spiders, whip spiders and vinegaroons.
Adult arachnids have eight legs attached to the cephalothorax. In some species the frontmost pair of legs has converted to a sensory function, while in others, different appendages can grow large enough to take on the appearance of extra pairs of legs.
Crustaceans (from Latin word "crustacea" meaning: "those with shells" or "crusted ones") are invertebrate animals that constitute one group of arthropods that are traditionally a part of the subphylum Crustacea (/krəˈsteɪʃə/), a large, diverse group of mainly aquatic arthropods including the more familiar decapods (shrimps, prawns, crabs, lobsters and crayfish), seed shrimps, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, opossum shrimps, amphipods and mantis shrimp. The crustacean group can be treated as a subphylum under the clade Mandibulata. It is now well accepted that the hexapods (insects and entognathans) emerged deep in the crustacean group, with the completed pan-group referred to as Pancrustacea. The three classes Cephalocarida, Branchiopoda and Remipedia are more closely related to the hexapods than they are to any of the other crustaceans (oligostracans and multicrustaceans).
The 67,000 described species range in size from Stygotantulus stocki at 0.1 mm (0.004 in), to the Japanese spider crab with a leg span of up to 3.8 m (12.5 ft) and a mass of 20 kg (44 lb). Like other arthropods, crustaceans have an exoskeleton, which they moult to grow. They are distinguished from other groups of arthropods, such as insects, myriapods and chelicerates, by the possession of biramous (two-parted) limbs, and by their larval forms, such as the nauplius stage of branchiopods and copepods.
A chela (/ˈkiːlə/) – also called a claw, nipper, or pincer – is a pincer-shaped organ at the end of certain limbs of some arthropods. The name comes from Ancient Greek χηλή, through Neo-Latin chela. The plural form is chelae. Legs bearing a chela are called chelipeds. Another name is claw because most chelae are curved and have a sharp point like a claw.
Chelae can be present at the tips of arthropod legs as well as their pedipalps. In some pseudoscorpions, the chelate pedipalps can be venomous. Chelae can be functionally overlapping with chelicerae, which specifically refers to appendages used as mouthparts in Chelicerata. In spiders those usually contain venom glands, but in other chelicerate allies the chelicerae can be chelate without venom glands.
Horseshoe crabs are arthropods of the family Limulidae and the only surviving xiphosurans. Despite their name, they are not crabs or even crustaceans; they are chelicerates, more closely related to arachnids like spiders, ticks, and scorpions. The body of a horseshoe crab is divided into three main parts: the cephalothorax, abdomen, and telson. The largest of these, the cephalothorax, houses most of the animal's eyes, limbs, and internal organs. It is also where the animal gets its name, as its shape somewhat resembles that of a horseshoe. Horseshoe crabs have been described as "living fossils", having changed little since they first appeared in the Triassic around 250 million years ago, and similar-looking fossil xiphosurans extend back to the Ordovician around 445 million years ago.
Only four species of horseshoe crab are extant today, the Atlantic horseshoe crab (Limulus polyphemus), native to the eastern coast of North and Central America, as well as the mangrove horseshoe crab (Carcinoscorpius rotundicauda), tri-spine horseshoe crab (Tachypleus tridentatus) and Indo-Pacific horseshoe crab (Tachypleus gigas), which are native to South, South East, and East Asia.
Eurypterids, often informally called sea scorpions, are a group of extinct marine arthropods that form the order Eurypterida. The earliest known eurypterids date to the Tremadocian stage of the Ordovician period, 480 million years ago. The group is likely to have appeared first during the Late Cambrian period. With approximately 250 species, the Eurypterida is the most diverse Paleozoic chelicerate order. Following their appearance during the Ordovician, eurypterids became major components of marine faunas during the Silurian, from which the majority of eurypterid species have been described. The Silurian genus Eurypterus accounts for more than 90% of all known eurypterid specimens. Though the group continued to diversify during the subsequent Devonian period, the eurypterids were heavily affected by the Late Devonian extinction event. They declined in numbers and diversity until becoming extinct during the Permian–Triassic extinction event (or sometime shortly before) 251.9 million years ago.
Although popularly called "sea scorpions", only the earliest eurypterids were marine; many later forms lived in brackish or fresh water, and they were not true scorpions. Some studies suggest that a dual respiratory system was present, which would have allowed for short periods of time in terrestrial environments. The name Eurypterida comes from Ancient Greek εὐρύς (eurús), meaning "wide, broad", and πτερόν (pterón), meaning "wing", referring to the pair of wide swimming appendages present in many members of the group.
The clade Mandibulata constitutes one of the major subdivisions of the phylum Arthropoda, alongside Chelicerata. Mandibulates include the crustaceans, myriapods (centipedes and millipedes, among others), and all true insects. The name "Mandibulata" refers to the mandibles, a modified pair of limbs used in food processing, the presence of which are characteristic of most members of the group.
The mandibulates are divided between the extant groups Myriapoda (millipedes and centipedes, among others) and Pancrustacea (including crustaceans and hexapods, the latter group containing insects). Molecular phylogenetic studies suggest that the living arthropods are related as shown in the cladogram below. Crustaceans do not form a monophyletic group as insects and other hexapods have evolved from within them.
Habeliida is an order of extinct arthropods that existed during the middle Cambrian. It is divided into two families, Habeliidae (monotypic, containing only Habelia), and Sanctacarididae (containing Sanctacaris, Utahcaris and Wisangocaris). They are thought to have been durophagous, with robust gnathobases (spined basal sections of limbs) used to shred hard-shelled organisms. Remains of trilobites have been found as stomach contents in Wisangocaris. Messorocaris has been suggested to be part of the order in some studies, but this is uncertain. They are suggested to be stem-group chelicerates, though they lack the chelicerae present in true chelicerates.
Cladogram after O’Flynn et al, 2023: