Charles Darwin

William Tecumseh Sherman Fitch III (born 1963) is an American evolutionary biologist and cognitive scientist at the University of Vienna (Vienna, Austria) where he is co-founder of the Department of Cognitive Biology. He was elected a Member of the National Academy of Sciences in 2025.

Fitch studies the biology and evolution of cognition and communication in humans and other animals, and in particular the evolution of speech, language and music. His work concentrates on comparative approaches as advocated by Charles Darwin (i.e., the study of homologous and analogous structures and processes in a wide range of species).

In biology, coevolution occurs when two or more species reciprocally affect each other's evolution through the process of natural selection. The term sometimes is used for two traits in the same species affecting each other's evolution, as well as gene-culture coevolution.

Charles Darwin mentioned evolutionary interactions between flowering plants and insects in On the Origin of Species (1859). He used use the similar word coadaptation and suggested how plants and insects could evolve through reciprocal evolutionary changes. Naturalists in the late 1800s studied other examples of how interactions among species could result in reciprocal evolutionary change. Beginning in the 1940s, plant pathologists developed breeding programs that were examples of human-induced coevolution. Development of new crop plant varieties that were resistant to some diseases favored rapid evolution in pathogen populations to overcome those plant defenses. That, in turn, required the development of yet new resistant crop plant varieties, producing an ongoing cycle of reciprocal evolution in crop plants and diseases that continues to this day.

Galápagos (Spanish pronunciation: [ɡaˈlapaɣos]) is a province of Ecuador in the country's Insular region, located approximately 1,000 km (620 mi) off the western coast of the mainland. The capital is Puerto Baquerizo Moreno.

The province administers the Galápagos Islands, a group of tiny volcanic islands that sit on the equator, famous for their unique biodiversity popularized by naturalist Charles Darwin and his theory of evolution.

The Galápagos Islands (Spanish: Islas Galápagos) are an archipelago of volcanic islands in the Eastern Pacific, located around the equator, 900 km (560 mi) west of the mainland of South America. They form the Galápagos Province of the Republic of Ecuador, with a population of slightly over 33,000 (2020). The province is divided into the cantons of San Cristóbal, Santa Cruz, and Isabela, the three most populated islands in the chain. The Galápagos are famous for their large number of endemic species, which were studied by Charles Darwin in the 1830s and inspired his theory of evolution by means of natural selection. All of these islands are protected as part of Ecuador's Galápagos National Park and Marine Reserve.

There is no firm evidence that Polynesians or the Indigenous peoples of South America reached the islands before their accidental discovery by Bishop Tomás de Berlanga in 1535. If some visitors did arrive, poor access to fresh water on the islands seems to have limited settlement. The Spanish Empire similarly ignored the islands, although during the Golden Age of Piracy various pirates used the Galápagos as a base for raiding Spanish shipping along the Peruvian coast. The goats and black and brown rats introduced during this period greatly damaged the existing ecosystems of several islands. British sailors were chiefly responsible for exploring and mapping the area. Darwin's voyage on HMS Beagle was part of an extensive British survey of the coasts of South America. Ecuador, which won its independence from Spain in 1822 and left Gran Colombia in 1830, formally occupied and claimed the islands on 12 February 1832 while the voyage was ongoing. José de Villamil, the founder of the Ecuadorian Navy, led the push to colonize and settle the islands, gradually supplanting the English names of the major islands with Spanish ones. The United States built the islands' first airport as a base to protect the western approaches of the Panama Canal in the 1930s. After World War II, its facilities were transferred to Ecuador. With the growing importance of ecotourism to the local economy, the airport modernized in the 2010s, using recycled materials for any expansion and shifting entirely to renewable energy sources to handle its roughly 300,000 visitors each year.

Animal culture can be defined as the ability of non-human animals to learn and transmit behaviors through processes of social or cultural learning.Culture is increasingly seen as a process, involving the social transmittance of behavior among peers and between generations. It can involve the transmission of novel behaviors or regional variations that are independent of genetic or ecological factors.

The existence of culture in non-humans has been a contentious subject, sometimes forcing researchers to rethink "what it is to be human".The notion of culture in other animals dates back to Aristotle in classical antiquity, and more recently to Charles Darwin, but the association of other animals' actions with the actual word 'culture' originated with Japanese primatologists' discoveries of socially-transmitted food behaviours in the 1940s. Evidence for animal culture is often based on studies of feeding behaviors, vocalizations, predator avoidance, mate selection, and migratory routes.

In biology, adaptation has three related meanings. Firstly, it is the dynamic evolutionary process of natural selection that fits organisms to their environment, enhancing their evolutionary fitness. Secondly, it is a state reached by the population during that process. Thirdly, it is a phenotypic trait or adaptive trait, with a functional role in each individual organism, that is maintained and has evolved through natural selection.

Historically, adaptation has been described from the time of the ancient Greek philosophers such as Empedocles and Aristotle. In 18th and 19th-century natural theology, adaptation was taken as evidence for the existence of a deity. Charles Darwin and Alfred Russel Wallace proposed instead that it was explained by natural selection.

Speciation is the evolutionary process by which populations evolve to become distinct species. The biologist Orator F. Cook coined the term in 1906 for cladogenesis, the splitting of lineages, as opposed to anagenesis, phyletic evolution within lineages. Charles Darwin was the first to describe the role of natural selection in speciation in his 1859 book On the Origin of Species. He also identified sexual selection as a likely mechanism, but found it problematic.

There are four geographic modes of speciation in nature, based on the extent to which speciating populations are isolated from one another: allopatric, peripatric, parapatric, and sympatric. Whether genetic drift is a minor or major contributor to speciation is the subject of much ongoing discussion.

Evolution is the change in the heritable characteristics of biological populations over successive generations. It occurs when evolutionary processes such as genetic drift and natural selection act on genetic variation, resulting in certain characteristics becoming more or less common within a population over successive generations. The process of evolution has given rise to biodiversity at every level of biological organisation.

The scientific theory of evolution by natural selection was conceived independently by two British naturalists, Charles Darwin and Alfred Russel Wallace, in the mid-19th century as an explanation for why organisms are adapted to their physical and biological environments. The theory was first set out in detail in Darwin's book On the Origin of Species. Evolution by natural selection is established by observable facts about living organisms: (1) more offspring are often produced than can possibly survive; (2) traits vary among individuals with respect to their morphology, physiology, and behaviour; (3) different traits confer different rates of survival and reproduction (differential fitness); and (4) traits can be passed from generation to generation (heritability of fitness). In successive generations, members of a population are therefore more likely to be replaced by the offspring of parents with favourable characteristics for that environment.

Adam Sedgwick FRS (/ˈsɛdʒwɪk/; 22 March 1785 – 27 January 1873) was a British geologist and Anglican priest, one of the founders of modern geology. He proposed the Cambrian and Devonian period of the geological timescale. Based on work which he did on Welsh rock strata, he proposed the Cambrian period in 1835, in a joint publication in which Roderick Murchison also proposed the Silurian period. Later in 1840, to resolve what later became known as the Great Devonian Controversy about rocks near the boundary between the Silurian and Carboniferous periods, he and Murchison proposed the Devonian period.

Though he had guided the young Charles Darwin in his early study of geology and continued to be on friendly terms, Sedgwick was an opponent of Darwin's theory of evolution by means of natural selection.

Paleoecology (also spelled palaeoecology) is the study of interactions between organisms and/or interactions between organisms and their environments across geologic timescales. As a discipline, paleoecology interacts with, depends on and informs a variety of fields including paleontology, ecology, climatology and biology.

Paleoecology emerged from the field of paleontology in the 1950s, though paleontologists have conducted paleoecological studies since the creation of paleontology in the 1700s and 1800s. Combining the investigative approach of searching for fossils with the theoretical approach of Charles Darwin and Alexander von Humboldt, paleoecology began as paleontologists began examining both the ancient organisms they discovered and the reconstructed environments in which they lived. Visual depictions of past marine and terrestrial communities have been considered an early form of paleoecology. The term "paleo-ecology" was coined by Frederic Clements in 1916.