Chalicotheriidae (from Ancient Greek χάλιξ (khálix), meaning "gravel", and θηρίον (theríon), meaning "beast") is an extinct family of herbivorous, perissodactyl mammals that lived in North America, Eurasia, and Africa from the Middle Eocene to the Early Pleistocene. They are often called chalicotheres, a term which is also applied to the broader grouping of Chalicotherioidea. They are noted for their unusual morphology compared to other ungulates, such as their clawed forelimbs. Members of the subfamily Chalicotheriinae developed elongated gorilla-like forelimbs that are thought to have been used to grasp vegetation. They are thought to have been browsers on foliage as well as possibly bark and fruit.
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Chalicotheriidae in the context of Odd-toed ungulate
Perissodactyla (/pəˌrɪsoʊˈdæktɪlə/, from Ancient Greek περισσός, perissós 'odd' and δάκτυλος, dáktylos 'finger, toe'), or odd-toed ungulates, is an order of ungulates. The order includes about 17 living species divided into three families: Equidae (horses, asses, and zebras), Rhinocerotidae (rhinoceroses), and Tapiridae (tapirs). They typically have reduced the weight-bearing toes to three or one of the five original toes, though tapirs retain four toes on their front feet. The nonweight-bearing toes are either present, absent, vestigial, or positioned posteriorly. By contrast, artiodactyls (even-toed ungulates) bear most of their weight equally on four or two (an even number) of the five toes: their third and fourth toes. Another difference between the two is that perissodactyls digest plant cellulose in their intestines, rather than in one or more stomach chambers as artiodactyls, with the exception of Suina, do.
The order was considerably more diverse in the past, with notable extinct groups including the brontotheres, palaeotheres, chalicotheres, and the paraceratheres, with the paraceratheres including the largest known land mammals to have ever existed.
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