Australopithecus sediba is an extinct species of australopithecine recovered from Malapa Cave, Cradle of Humankind, South Africa. It is known from a partial juvenile skeleton, the holotype MH1, and a partial adult female skeleton, the paratype MH2. They date to about 1.98 million years ago in the Early Pleistocene, and coexisted with Paranthropus robustus and Homo ergaster / Homo erectus. Malapa Cave may have been a natural death trap, the base of a long vertical shaft which creatures could accidentally fall into. A. sediba was initially described as being a potential human ancestor, and perhaps the progenitor of Homo, but this is contested and it could also represent a late-surviving population or sister species of A. africanus which had earlier inhabited the area.
MH1 has a brain volume of about 350–440 cc, similar to other australopithecines. The face of MH1 is strikingly similar to Homo instead of other australopithecines, with a less pronounced brow ridge, cheek bones, and prognathism (the amount the face juts out), and there is evidence of a slight chin. However, such characteristics could be due to juvenility and lost with maturity. The teeth are quite small for an australopithecine. MH1 is estimated at 130 cm (4 ft 3 in) tall, which would equate to an adult height of 150–156 cm (4 ft 11 in – 5 ft 1 in). MH1 and MH2 were estimated to have been about the same weight at 30–36 kg (66–79 lb). Like other australopithecines, A. sediba is thought to have had a narrow and apelike upper chest, but a broad and humanlike lower chest. Like other australopithecines, the arm anatomy seems to suggest a degree of climbing and arboreal behaviour. The pelvis indicates A. sediba was capable of a humanlike stride, but the foot points to a peculiar gait not demonstrated in any other hominin involving hyperpronation of the ankle, and resultantly rotating the leg inwards while pushing off. This suite of adaptations may represent a compromise between habitual bipedalism and arboreality.
